{
    "glytoucan_ac": "G02686MD",
    "number_monosaccharides": 22,
    "number_monosaccharides_suffix": "",
    "mass": 4413.58,
    "mass_pme": 5478.77,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "no",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "no",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b1-D-Neup5Ac-(2\u21926)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)[\u03b1-D-Neup5Ac-(2\u21926)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)]-\u03b1-D-Manp-(1\u21923)[\u03b1-D-Neup5Ac-(2\u21926)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)[\u03b1-D-Neup5Ac-(2\u21926)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21926)]-\u03b1-D-Manp-(1\u21926)]-\u03b2-D-Manp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)[\u03b1-L-Fucp-(1\u21926)]-?-D-GlcNAc-ol",
    "wurcs": "WURCS=2.0/7,22,21/[h2122h_2*NCC/3=O][a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O][a1221m-1a_1-5]/1-2-3-4-2-5-6-2-5-6-4-2-5-2-5-6-2-5-2-5-6-7/a4-b1_a6-v1_b4-c1_c3-d1_c6-k1_d2-e1_d4-h1_e4-f1_f6-g2_h4-i1_i6-j2_k2-l1_k6-q1_l4-m1_m4-n1_n4-o1_o6-p2_q4-r1_r4-s1_s4-t1_t6-u2",
    "glycoct": "RES 1b:o-dglc-HEX-0:0|1:aldi 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 11s:n-acetyl 12b:b-dglc-HEX-1:5 13s:n-acetyl 14b:b-dgal-HEX-1:5 15b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 16s:n-acetyl 17b:a-dman-HEX-1:5 18b:b-dglc-HEX-1:5 19s:n-acetyl 20b:b-dgal-HEX-1:5 21b:b-dglc-HEX-1:5 22s:n-acetyl 23b:b-dgal-HEX-1:5 24b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 25s:n-acetyl 26b:b-dglc-HEX-1:5 27s:n-acetyl 28b:b-dgal-HEX-1:5 29b:b-dglc-HEX-1:5 30s:n-acetyl 31b:b-dgal-HEX-1:5 32b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 33s:n-acetyl 34b:a-lgal-HEX-1:5|6:d LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:9o(6+2)10d 10:10d(5+1)11n 11:6o(4+1)12d 12:12d(2+1)13n 13:12o(4+1)14d 14:14o(6+2)15d 15:15d(5+1)16n 16:5o(6+1)17d 17:17o(2+1)18d 18:18d(2+1)19n 19:18o(4+1)20d 20:20o(4+1)21d 21:21d(2+1)22n 22:21o(4+1)23d 23:23o(6+2)24d 24:24d(5+1)25n 25:17o(6+1)26d 26:26d(2+1)27n 27:26o(4+1)28d 28:28o(4+1)29d 29:29d(2+1)30n 30:29o(4+1)31d 31:31o(6+2)32d 32:32d(5+1)33n 33:1o(6+1)34d",
    "inchi": "InChI=1S/C168H276N12O122/c1-40-90(217)109(236)117(244)151(268-40)261-33-62(216)126(91(218)53(18-181)169-41(2)196)285-145-84(175-47(8)202)103(230)133(69(29-192)270-145)295-158-125(252)141(296-160-143(298-150-89(180-52(13)207)108(235)131(67(27-190)275-150)292-155-121(248)113(240)99(226)78(283-155)39-267-168(164(259)260)17-57(211)82(173-45(6)200)140(302-168)95(222)61(215)22-185)124(251)136(72(32-195)278-160)288-148-87(178-50(11)205)106(233)130(66(26-189)273-148)291-154-120(247)112(239)98(225)77(282-154)38-266-167(163(257)258)16-56(210)81(172-44(5)199)139(301-167)94(221)60(214)21-184)101(228)74(279-158)35-263-159-142(297-149-88(179-51(12)206)107(234)132(68(28-191)274-149)294-157-123(250)116(243)135(71(31-194)277-157)287-147-86(177-49(10)204)105(232)129(65(25-188)272-147)290-153-119(246)111(238)97(224)76(281-153)37-265-166(162(255)256)15-55(209)80(171-43(4)198)138(300-166)93(220)59(213)20-183)114(241)100(227)73(284-159)34-262-144-83(174-46(7)201)102(229)127(63(23-186)269-144)293-156-122(249)115(242)134(70(30-193)276-156)286-146-85(176-48(9)203)104(231)128(64(24-187)271-146)289-152-118(245)110(237)96(223)75(280-152)36-264-165(161(253)254)14-54(208)79(170-42(3)197)137(299-165)92(219)58(212)19-182/h40,53-160,181-195,208-252H,14-39H2,1-13H3,(H,169,196)(H,170,197)(H,171,198)(H,172,199)(H,173,200)(H,174,201)(H,175,202)(H,176,203)(H,177,204)(H,178,205)(H,179,206)(H,180,207)(H,253,254)(H,255,256)(H,257,258)(H,259,260)/t40-,53-,54-,55-,56-,57-,58+,59+,60+,61+,62+,63+,64+,65+,66+,67+,68+,69+,70+,71+,72+,73+,74+,75+,76+,77+,78+,79+,80+,81+,82+,83+,84+,85+,86+,87+,88+,89+,90+,91+,92+,93+,94+,95+,96-,97-,98-,99-,100+,101+,102+,103+,104+,105+,106+,107+,108+,109+,110-,111-,112-,113-,114-,115+,116+,117-,118+,119+,120+,121+,122+,123+,124-,125-,126+,127+,128+,129+,130+,131+,132+,133+,134-,135-,136+,137+,138+,139+,140+,141-,142-,143-,144+,145-,146-,147-,148-,149-,150-,151+,152-,153-,154-,155-,156-,157-,158-,159-,160+,165+,166+,167+,168+/m0/s1",
    "glycam": "",
    "smiles_isomeric": "",
    "inchi_key": {
        "key": "QKJFIQZSAZPEKF-FRSINXOGSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/QKJFIQZSAZPEKF-FRSINXOGSA-N"
    },
    "byonic": "",
    "gwb": "redEnd--?D-GlcNAc,o(--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p(--2b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuAc,p)--4b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuAc,p)--6a1D-Man,p(--2b1D-GlcNAc,p--4b1D-Gal,p--4b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuAc,p)--6b1D-GlcNAc,p--4b1D-Gal,p--4b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuAc,p)--6a1L-Fuc,p$MONO,Und,0,0,redEnd",
    "species": [],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Alditol-reduced"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Complex",
                "url": ""
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Core-fucosylated"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Tetraantennary"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10469-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02686MD",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q10469",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02686MD",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02686MD"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921V5-1",
            "xref_key": "glycan_xref_sandbox",
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