{
    "glytoucan_ac": "G08520NM",
    "number_monosaccharides": 8,
    "number_monosaccharides_suffix": "",
    "mass": 1437.51,
    "mass_pme": 1801.92,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "yes",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b2-D-GlcpNAc-(1\u21922)-\u03b1-D-Manp-(1\u21923)[\u03b1-D-Manp-(1\u21923)[\u03b1-D-Manp-(1\u21926)]-\u03b1-D-Manp-(1\u21926)]-\u03b2-D-Manp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)-\u03b2-D-GlcpNAc-(1\u2192",
    "wurcs": "WURCS=2.0/3,8,7/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5]/1-1-2-3-1-3-3-3/a4-b1_b4-c1_c3-d1_c6-f1_d2-e1_f3-g1_f6-h1",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:a-dman-HEX-1:5 10b:a-dman-HEX-1:5 11b:a-dman-HEX-1:5 LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:5o(6+1)9d 9:9o(3+1)10d 10:9o(6+1)11d",
    "inchi": "InChI=1S/C54H91N3O41/c1-12(64)55-23-33(74)42(19(8-62)86-47(23)83)94-49-25(57-14(3)66)34(75)43(20(9-63)91-49)95-53-41(82)45(97-54-46(37(78)29(70)18(7-61)90-54)98-48-24(56-13(2)65)32(73)26(67)15(4-58)87-48)31(72)22(93-53)11-85-51-40(81)44(96-52-39(80)36(77)28(69)17(6-60)89-52)30(71)21(92-51)10-84-50-38(79)35(76)27(68)16(5-59)88-50/h15-54,58-63,67-83H,4-11H2,1-3H3,(H,55,64)(H,56,65)(H,57,66)/t15-,16-,17-,18-,19-,20-,21-,22-,23-,24-,25-,26-,27-,28-,29-,30-,31-,32-,33-,34-,35+,36+,37+,38+,39+,40+,41+,42-,43-,44+,45+,46+,47-,48+,49+,50+,51+,52-,53+,54-/m1/s1",
    "glycam": "DManpa1-3[DManpa1-6]DManpa1-6[DGlcpNAcb1-2DManpa1-3]DManpb1-4DGlcpNAcb1-4DGlcpNAcb1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H]([C@@H]([C@H](O[C@H]1O)CO)O[C@H]2[C@@H]([C@H]([C@@H]([C@H](O2)CO)O[C@H]3[C@H]([C@H]([C@@H]([C@H](O3)CO[C@@H]4[C@H]([C@H]([C@@H]([C@H](O4)CO[C@@H]5[C@H]([C@H]([C@@H]([C@H](O5)CO)O)O)O)O)O[C@@H]6[C@H]([C@H]([C@@H]([C@H](O6)CO)O)O)O)O)O)O[C@@H]7[C@H]([C@H]([C@@H]([C@H](O7)CO)O)O)O[C@H]8[C@@H]([C@H]([C@@H]([C@H](O8)CO)O)O)NC(=O)C)O)O)NC(=O)C)O",
    "inchi_key": {
        "key": "MRIXKRYHOZQBAW-ABMGQQJYSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/MRIXKRYHOZQBAW-ABMGQQJYSA-N"
    },
    "byonic": "HexNAc(3)Hex(5) % 1437.51280002",
    "gwb": "freeEnd--1b1D-GlcNAc,p--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p--2b1D-GlcNAc,p)--6a1D-Man,p(--3a1D-Man,p)--6a1D-Man,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9913,
            "name": "Bos taurus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3141",
                    "url": "https://glyconnect.expasy.org/browser/structures/3141"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G08520NM",
                    "url": "https://glycosmos.org/glycans/show/G08520NM"
                },
                {
                    "database": "PubMed",
                    "id": "447724",
                    "url": "https://glygen.org/publication/PubMed/447724"
                }
            ],
            "common_name": "Bovine",
            "glygen_name": "Bovine",
            "reference_species": "Bos taurus [9913]"
        },
        {
            "taxid": 9031,
            "name": "Gallus gallus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3141",
                    "url": "https://glyconnect.expasy.org/browser/structures/3141"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G08520NM",
                    "url": "https://glycosmos.org/glycans/show/G08520NM"
                },
                {
                    "database": "PubMed",
                    "id": "10833030",
                    "url": "https://glygen.org/publication/PubMed/10833030"
                }
            ],
            "common_name": "Chicken",
            "glygen_name": "Chicken",
            "reference_species": "Gallus gallus [9031]"
        },
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "11708",
                    "url": "https://glyconnect.expasy.org/browser/structures/11708"
                },
                {
                    "database": "GlyConnect",
                    "id": "3141",
                    "url": "https://glyconnect.expasy.org/browser/structures/3141"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G08520NM",
                    "url": "https://glycosmos.org/glycans/show/G08520NM"
                },
                {
                    "database": "PubMed",
                    "id": "10207177",
                    "url": "https://glygen.org/publication/PubMed/10207177"
                },
                {
                    "database": "PubMed",
                    "id": "11469797",
                    "url": "https://glygen.org/publication/PubMed/11469797"
                },
                {
                    "database": "PubMed",
                    "id": "24706782",
                    "url": "https://glygen.org/publication/PubMed/24706782"
                },
                {
                    "database": "PubMed",
                    "id": "25802287",
                    "url": "https://glygen.org/publication/PubMed/25802287"
                },
                {
                    "database": "PubMed",
                    "id": "26510530",
                    "url": "https://glygen.org/publication/PubMed/26510530"
                },
                {
                    "database": "PubMed",
                    "id": "32643525",
                    "url": "https://glygen.org/publication/PubMed/32643525"
                },
                {
                    "database": "PubMed",
                    "id": "33139572",
                    "url": "https://glygen.org/publication/PubMed/33139572"
                },
                {
                    "database": "PubMed",
                    "id": "7514386",
                    "url": "https://glygen.org/publication/PubMed/7514386"
                },
                {
                    "database": "PubMed",
                    "id": "7881178",
                    "url": "https://glygen.org/publication/PubMed/7881178"
                },
                {
                    "database": "UniCarbKB",
                    "id": "GLY_000341",
                    "url": "http://data.glygen.org/GLY_000341"
                }
            ],
            "common_name": "Human",
            "glygen_name": "Human",
            "reference_species": "Homo sapiens [9606]"
        },
        {
            "taxid": 10116,
            "name": "Rattus norvegicus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3141",
                    "url": "https://glyconnect.expasy.org/browser/structures/3141"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G08520NM",
                    "url": "https://glycosmos.org/glycans/show/G08520NM"
                },
                {
                    "database": "PubMed",
                    "id": "10207177",
                    "url": "https://glygen.org/publication/PubMed/10207177"
                },
                {
                    "database": "PubMed",
                    "id": "24090084",
                    "url": "https://glygen.org/publication/PubMed/24090084"
                },
                {
                    "database": "UniCarbKB",
                    "id": "GLY_000341",
                    "url": "http://data.glygen.org/GLY_000341"
                }
            ],
            "common_name": "Rat",
            "glygen_name": "Rat",
            "reference_species": "Rattus norvegicus [10116]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Hybrid",
                "url": ""
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
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            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22",
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            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
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                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
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        },
        {
            "uniprot_canonical_ac": "Q8K2A8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "Dol-P-Man:Man(5)GlcNAc(2)-PP-Dol alpha-1,3-mannosyltransferase",
            "gene": "Alg3",
            "gene_link": "https://www.uniprot.org/uniprot/Q8K2A8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
                }
            ]
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        {
            "uniprot_canonical_ac": "Q92685-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08520NM",
            "protein_name": "Dol-P-Man:Man(5)GlcNAc(2)-PP-Dol alpha-1,3-mannosyltransferase",
            "gene": "ALG3",
            "gene_link": "https://www.uniprot.org/uniprot/Q92685",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08520NM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08520NM"
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            ]
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    "crossref": [
        {
            "id": "91858075",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/91858075",
            "database": "PubChem Compound",
            "categories": [
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            ]
        },
        {
            "id": "252291399",
            "url": "https://pubchem.ncbi.nlm.nih.gov/substance/252291399",
            "database": "PubChem Substance",
            "categories": [
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            ]
        },
        {
            "id": "2545",
            "url": "",
            "database": "UniCarbKB",
            "categories": [
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            ]
        },
        {
            "id": "G00013",
            "url": "https://www.genome.jp/dbget-bin/www_bget?gl:G00013",
            "database": "KEGG Glycan",
            "categories": [
                "Enzyme and Pathway"
            ]
        },
        {
            "id": "23657",
            "url": "",
            "database": "Glycosciences.de",
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            ]
        },
        {
            "id": "2384",
            "url": "",
            "database": "Glycosciences.de",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "11708",
            "url": "https://glyconnect.expasy.org/browser/structures/11708",
            "database": "GlyConnect",
            "categories": [
                "Glycoproteomics",
                "Glycomics"
            ]
        },
        {
            "id": "3141",
            "url": "https://glyconnect.expasy.org/browser/structures/3141",
            "database": "GlyConnect",
            "categories": [
                "Glycoproteomics",
                "Glycomics"
            ]
        },
        {
            "id": "11938",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+11938",
            "database": "CarbBank",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "29587",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+29587",
            "database": "CarbBank",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "31282",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+31282",
            "database": "CarbBank",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "33013",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+33013",
            "database": "CarbBank",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "35726",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+35726",
            "database": "CarbBank",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "38414",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+38414",
            "database": "CarbBank",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "38483",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+38483",
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