{
    "glytoucan_ac": "G06953DR",
    "number_monosaccharides": 7,
    "number_monosaccharides_suffix": "",
    "mass": 1492.52,
    "mass_pme": 1870.94,
    "tool_support": {
        "gnome_glygen_oglycans": "yes",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "yes",
        "pdb": "yes",
        "gnome_glygen_nglycans": "no",
        "gnome_glycotree_nglycans": "no"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b1-D-Neup5Ac-(2\u21923)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)[\u03b1-D-Neup5Ac-(2\u21923)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21926)]-\u03b1-D-Manp-(1\u2192",
    "wurcs": "WURCS=2.0/4,7,6/[a1122h-1a_1-5][a2122h-1b_1-5_2*NCC/3=O][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O]/1-2-3-4-2-3-4/a2-b1_a6-e1_b4-c1_c3-d2_e4-f1_f3-g2",
    "glycoct": "RES 1b:a-dman-HEX-1:5 2b:b-dglc-HEX-1:5 3s:n-acetyl 4b:b-dgal-HEX-1:5 5b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 6s:n-acetyl 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 11s:n-acetyl LIN 1:1o(2+1)2d 2:2d(2+1)3n 3:2o(4+1)4d 4:4o(3+2)5d 5:5d(5+1)6n 6:1o(6+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:9o(3+2)10d 10:10d(5+1)11n",
    "inchi": "InChI=1S/C56H92N4O42/c1-14(67)57-27-18(71)5-55(53(86)87,99-43(27)31(75)20(73)7-61)101-45-34(78)22(9-63)92-51(39(45)83)96-41-24(11-65)94-49(29(36(41)80)59-16(3)69)90-13-26-33(77)38(82)47(48(85)91-26)98-50-30(60-17(4)70)37(81)42(25(12-66)95-50)97-52-40(84)46(35(79)23(10-64)93-52)102-56(54(88)89)6-19(72)28(58-15(2)68)44(100-56)32(76)21(74)8-62/h18-52,61-66,71-85H,5-13H2,1-4H3,(H,57,67)(H,58,68)(H,59,69)(H,60,70)(H,86,87)(H,88,89)/t18-,19-,20+,21+,22+,23+,24+,25+,26+,27+,28+,29+,30+,31+,32+,33+,34-,35-,36+,37+,38-,39+,40+,41+,42+,43+,44+,45-,46-,47-,48-,49+,50-,51-,52-,55-,56-/m0/s1",
    "glycam": "DNeup5Aca2-3DGalpb1-4DGlcpNAcb1-2[DNeup5Aca2-3DGalpb1-4DGlcpNAcb1-6]DManpa1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H](C[C@@](O[C@H]1[C@@H]([C@@H](CO)O)O)(C(=O)O)O[C@H]2[C@H]([C@H](O[C@H]([C@@H]2O)O[C@@H]3[C@H](O[C@H]([C@@H]([C@H]3O)NC(=O)C)OC[C@@H]4[C@H]([C@@H]([C@@H]([C@H](O4)O)O[C@H]5[C@@H]([C@H]([C@@H]([C@H](O5)CO)O[C@H]6[C@@H]([C@H]([C@H]([C@H](O6)CO)O)O[C@@]7(C[C@@H]([C@H]([C@@H](O7)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)NC(=O)C)O)O)CO)CO)O)O",
    "inchi_key": {
        "key": "DLOQVCBLOYEWSC-IZAVBOCFSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/DLOQVCBLOYEWSC-IZAVBOCFSA-N"
    },
    "byonic": "HexNAc(2)Hex(3)NeuAc(2) % 1492.51861368",
    "gwb": "freeEnd--1a1D-Man,p(--2b1D-GlcNAc,p--4b1D-Gal,p--3a2D-NeuAc,p)--6b1D-GlcNAc,p--4b1D-Gal,p--3a2D-NeuAc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "11430",
                    "url": "https://glyconnect.expasy.org/browser/structures/11430"
                },
                {
                    "database": "GlyConnect",
                    "id": "40",
                    "url": "https://glyconnect.expasy.org/browser/structures/40"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G06953DR",
                    "url": "https://glycosmos.org/glycans/show/G06953DR"
                },
                {
                    "database": "PubMed",
                    "id": "34191522",
                    "url": "https://glygen.org/publication/PubMed/34191522"
                }
            ],
            "common_name": "Human",
            "glygen_name": "Human",
            "reference_species": "Homo sapiens [9606]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "O-linked",
                "url": ""
            },
            "subtype": {
                "name": "O-mannose core"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q8BGQ4-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Protein O-mannosyl-transferase 2",
            "gene": "Pomt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BGQ4",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8R2R1-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Protein O-mannosyl-transferase 1",
            "gene": "Pomt1",
            "gene_link": "https://www.uniprot.org/uniprot/Q8R2R1",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9UKY4-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Protein O-mannosyl-transferase 2",
            "gene": "POMT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9UKY4",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Y6A1-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Protein O-mannosyl-transferase 1",
            "gene": "POMT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Y6A1",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8WZA1-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Protein O-linked-mannose beta-1,2-N-acetylglucosaminyltransferase 1",
            "gene": "POMGNT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q8WZA1",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91X88-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Protein O-linked-mannose beta-1,2-N-acetylglucosaminyltransferase 1",
            "gene": "Pomgnt1",
            "gene_link": "https://www.uniprot.org/uniprot/Q91X88",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q3V5L5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Alpha-1,6-mannosylglycoprotein 6-beta-N-acetylglucosaminyltransferase B",
            "gene": "MGAT5B",
            "gene_link": "https://www.uniprot.org/uniprot/Q3V5L5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q765H6-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Alpha-1,6-mannosylglycoprotein 6-beta-N-acetylglucosaminyltransferase B",
            "gene": "Mgat5b",
            "gene_link": "https://www.uniprot.org/uniprot/Q765H6",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O43286-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 5",
            "gene": "B4GALT5",
            "gene_link": "https://www.uniprot.org/uniprot/O43286",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60513-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 4",
            "gene": "B4GALT4",
            "gene_link": "https://www.uniprot.org/uniprot/O60513",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9JJ04-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 4",
            "gene": "B4galt4",
            "gene_link": "https://www.uniprot.org/uniprot/Q9JJ04",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9JMK0-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 5",
            "gene": "B4galt5",
            "gene_link": "https://www.uniprot.org/uniprot/Q9JMK0",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P54751-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 1",
            "gene": "St3gal1",
            "gene_link": "https://www.uniprot.org/uniprot/P54751",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q11201-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G06953DR",
            "protein_name": "CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 1",
            "gene": "ST3GAL1",
            "gene_link": "https://www.uniprot.org/uniprot/Q11201",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G06953DR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G06953DR"
                }
            ]
        }
    ],
    "crossref": [
        {
            "id": "146016808",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/146016808",
            "database": "PubChem Compound",
            "categories": [
                "Chemical"
            ]
        },
        {
            "id": "405039480",
            "url": "https://pubchem.ncbi.nlm.nih.gov/substance/405039480",
            "database": "PubChem Substance",
            "categories": [
                "Chemical"
            ]
        },
        {
            "id": "11430",
            "url": "https://glyconnect.expasy.org/browser/structures/11430",
            "database": "GlyConnect",
            "categories": [
                "Glycoproteomics",
                "Glycomics"
            ]
        },
        {
            "id": "40",
            "url": "https://glyconnect.expasy.org/browser/structures/40",
            "database": "GlyConnect",
            "categories": [
                "Glycoproteomics",
                "Glycomics"
            ]
        },
        {
            "id": "157265",
            "url": "https://www.ebi.ac.uk/chebi/searchId.do?chebiId=157265",
            "database": "ChEBI",
            "categories": []
        },
        {
            "id": "G06953DR",
            "url": "https://glytoucan.org/Structures/Glycans/G06953DR",
            "database": "GlyTouCan",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "G06953DR",
            "url": "https://sandbox.glyomics.org/explore.html?G06953DR",
            "database": "SandBox",
            "categories": [
                "Enzyme and Pathway"
            ]
        },
        {
            "id": "G06953DR",
            "url": "https://gnome.glyomics.org/restrictions/GlyGen.StructureBrowser.html?focus=G06953DR",
            "database": "GNOme",
            "categories": [
                "Ontology and Dictionary",
                "Glycomics"
            ]
        },
        {
            "id": "G06953DR",
            "url": "https://glycosmos.org/glycans/show/G06953DR",
            "database": "GlyCosmos",
            "categories": [
                "Glycoproteomics",
                "Glycomics"
            ]
        },
        {
            "id": "GSD000001",
            "url": "https://wiki.glygen.org/Glycan_structure_dictionary#GSD000001",
            "database": "Glycan Structure Dictionary",
            "categories": [
                "Ontology and Dictionary"
            ]
        }
    ],
    "glycoprotein": [],
    "publication": [
        {
            "title": "The O-Glycome of Human Nigrostriatal Tissue and Its Alteration in Parkinson's Disease.",
            "journal": "Journal of proteome research",
            "date": "2021",
            "authors": "Wilkinson H, Thomsson KA, Rebelo AL, Hilliard M, Pandit A, Rudd PM, Karlsson NG, Saldova R",
            "evidence": [
                {
                    "database": "GlyTouCan",
                    "id": "G06953DR",
                    "url": "https://glytoucan.org/Structures/Glycans/G06953DR"
                },
                {
                    "database": "GlyConnect",
                    "id": "11430",
                    "url": "https://glyconnect.expasy.org/browser/structures/11430"
                }
            ],
            "reference": [
                {
                    "type": "PubMed",
                    "id": "34191522",
                    "url": "https://glygen.org/publication/PubMed/34191522"
                }
            ]
        },
        {
            "title": "High prevalence of 2-mono- and 2,6-di-substituted manol-terminating sequences among O-glycans released from brain glycopeptides by reductive alkaline hydrolysis.",
            "journal": "European journal of biochemistry",
            "date": "1999",
            "authors": "Chai W, Yuen CT, Kogelberg H, Carruthers RA, Margolis RU, Feizi T, Lawson AM",
            "evidence": [
                {
                    "database": "Glycan Motif",
                    "id": "GGM.000056",
                    "url": "https://glycomotif.glyomics.org/glycomotif/GGM.000056"
                }
            ],
            "reference": [
                {
                    "type": "PubMed",
                    "id": "10469154",
                    "url": "https://glygen.org/publication/PubMed/10469154"
                }
            ]
        },
        {
            "title": "The repertoire of glycan determinants in the human glycome.",
            "journal": "Molecular bioSystems",
            "date": "2009",
            "authors": "Cummings RD",
            "evidence": [
                {
                    "database": "Glycan Motif",
                    "id": "GGM.000056",
                    "url": "https://glycomotif.glyomics.org/glycomotif/GGM.000056"
                }
            ],
            "reference": [
                {
                    "type": "PubMed",
                    "id": "19756298",
                    "url": "https://glygen.org/publication/PubMed/19756298"
                }
            ]
        }
    ],
    "residues": [
        {
            "id": "OC4",
            "name": "O-glycan_core_x-D-Manx_C4",
            "attachedby": "rxn.Q8BGQ4-1",
            "detachedby": "",
            "parentid": "0"
        },
        {
            "id": "OC4",
            "name": "O-glycan_core_x-D-Manx_C4",
            "attachedby": "rxn.Q8R2R1-1",
            "detachedby": "",
            "parentid": "0"
        },
        {
            "id": "OC4",
            "name": "O-glycan_core_x-D-Manx_C4",
            "attachedby": "rxn.Q9UKY4-1",
            "detachedby": "",
            "parentid": "0"
        },
        {
            "id": "OC4",
            "name": "O-glycan_core_x-D-Manx_C4",
            "attachedby": "rxn.Q9Y6A1-1",
            "detachedby": "",
            "parentid": "0"
        },
        {
            "id": "O806",
            "name": "O-glycan_b-D-GlcpNAc_806",
            "attachedby": "rxn.Q8WZA1-1",
            "detachedby": "",
            "parentid": "OC4"
        },
        {
            "id": "O806",
            "name": "O-glycan_b-D-GlcpNAc_806",
            "attachedby": "rxn.Q91X88-1",
            "detachedby": "",
            "parentid": "OC4"
        },
        {
            "id": "O807",
            "name": "O-glycan_b-D-GlcpNAc_807",
            "attachedby": "rxn.Q3V5L5-1",
            "detachedby": "",
            "parentid": "OC4"
        },
        {
            "id": "O807",
            "name": "O-glycan_b-D-GlcpNAc_807",
            "attachedby": "rxn.Q765H6-1",
            "detachedby": "",
            "parentid": "OC4"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.O43286-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.O60512-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.O60513-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.O60909-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.P15291-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.P15535-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.Q91YY2-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.Q9JJ04-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.Q9JMK0-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O808",
            "name": "O-glycan_b-D-Galp_808",
            "attachedby": "rxn.Q9Z2Y2-1",
            "detachedby": "",
            "parentid": "O806"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.O43286-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.O60512-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.O60513-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.O60909-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.P15291-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.P15535-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.Q91YY2-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.Q9JJ04-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.Q9JMK0-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O809",
            "name": "O-glycan_b-D-Galp_809",
            "attachedby": "rxn.Q9Z2Y2-1",
            "detachedby": "",
            "parentid": "O807"
        },
        {
            "id": "O812",
            "name": "O-glycan_a-D-NeupNAc_812",
            "attachedby": "rxn.P54751-1",
            "detachedby": "",
            "parentid": "O808"
        },
        {
            "id": "O812",
            "name": "O-glycan_a-D-NeupNAc_812",
            "attachedby": "rxn.Q11201-1",
            "detachedby": "",
            "parentid": "O808"
        },
        {
            "id": "O813",
            "name": "O-glycan_a-D-NeupNAc_813",
            "attachedby": "rxn.P54751-1",
            "detachedby": "",
            "parentid": "O809"
        },
        {
            "id": "O813",
            "name": "O-glycan_a-D-NeupNAc_813",
            "attachedby": "rxn.Q11201-1",
            "detachedby": "",
            "parentid": "O809"
        }
    ],
    "composition": [
        {
            "name": "6-Deoxy-Hexose",
            "residue": "dhex",
            "count": 0,
            "cid": "840",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
        },
        {
            "name": "Hexose",
            "residue": "hex",
            "count": 3,
            "cid": "206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206"
        },
        {
            "name": "N-Acetylhexosamine",
            "residue": "hexnac",
            "count": 2,
            "cid": "899",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
        },
        {
            "name": "Hexuronic Acid",
            "residue": "hexa",
            "count": 0,
            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
        },
        {
            "name": "2-Amino-2-Deoxy-Hexose",
            "residue": "hexn",
            "count": 0,
            "cid": "739",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739"
        },
        {
            "name": "N-Acetyl-Neuraminic Acid",
            "residue": "neuac",
            "count": 2,
            "cid": "439197",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
        },
        {
            "name": "N-Glycolyl-Neuraminic Acid",
            "residue": "neugc",
            "count": 0,
            "cid": "440001",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
        },
        {
            "name": "Phosphate",
            "residue": "p",
            "count": 0,
            "cid": "1061",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
        },
        {
            "name": "Pentose",
            "residue": "pent",
            "count": 0,
            "cid": "229",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
        },
        {
            "name": "Sulfate",
            "residue": "s",
            "count": 0,
            "cid": "1117",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
        },
        {
            "name": "Other",
            "residue": "other",
            "count": 0
        }
    ],
    "composition_expanded": [
        {
            "name": "Alditol",
            "residue": "aldi",
            "count": 0
        },
        {
            "name": "6-Deoxy-Hexose",
            "residue": "dhex",
            "count": 0,
            "cid": "840",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
        },
        {
            "name": "6-Deoxy-Hexitol",
            "residue": "dhex+aldi",
            "count": 0
        },
        {
            "name": "L-Fucose",
            "residue": "fuc",
            "count": 0,
            "cid": "17106",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/17106"
        },
        {
            "name": "L-Fucitol",
            "residue": "fuc+aldi",
            "count": 0,
            "cid": "445724",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/445724"
        },
        {
            "name": "D-Galactose",
            "residue": "gal",
            "count": 2,
            "cid": "6036",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6036"
        },
        {
            "name": "D-Galactitol",
            "residue": "gal+aldi",
            "count": 0,
            "cid": "11850",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/11850"
        },
        {
            "name": "N-Acetyl-D-galactosamine",
            "residue": "galnac",
            "count": 0,
            "cid": "35717",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/35717"
        },
        {
            "name": "N-Acetylgalactosaminitol",
            "residue": "galnac+aldi",
            "count": 0,
            "cid": "165880",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/165880"
        },
        {
            "name": "D-Galacturonic Acid",
            "residue": "gala",
            "count": 0,
            "cid": "439215",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439215"
        },
        {
            "name": "D-Glucose",
            "residue": "glc",
            "count": 0,
            "cid": "5793",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/5793"
        },
        {
            "name": "D-Glucitol",
            "residue": "glc+aldi",
            "count": 0,
            "cid": "5780",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/5780"
        },
        {
            "name": "D-Glucuronic acid",
            "residue": "glca",
            "count": 0,
            "cid": "94715",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/94715"
        },
        {
            "name": "D-Glucosamine",
            "residue": "glcn",
            "count": 0,
            "cid": "439213",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439213"
        },
        {
            "name": "N-Acetyl-D-Glucosamine",
            "residue": "glcnac",
            "count": 2,
            "cid": "439174",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439174"
        },
        {
            "name": "N-Acetyl-D-glucosaminitol",
            "residue": "glcnac+aldi",
            "count": 0,
            "cid": "165206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/165206"
        },
        {
            "name": "Hexose",
            "residue": "hex",
            "count": 3,
            "cid": "206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206"
        },
        {
            "name": "N-Acetylhexosamine",
            "residue": "hexnac",
            "count": 2,
            "cid": "899",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
        },
        {
            "name": "Hexuronic Acid",
            "residue": "hexa",
            "count": 0,
            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
        },
        {
            "name": "2-Amino-2-Deoxy-Hexose",
            "residue": "hexn",
            "count": 0,
            "cid": "739",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739"
        },
        {
            "name": "Hexitol",
            "residue": "hex+aldi",
            "count": 0,
            "cid": "453",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/453"
        },
        {
            "name": "N-Acetylhexosaminitol",
            "residue": "hexnac+aldi",
            "count": 0
        },
        {
            "name": "L-Iduronic acid",
            "residue": "idoa",
            "count": 0,
            "cid": "441039",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/441039"
        },
        {
            "name": "D-Mannose",
            "residue": "man",
            "count": 1,
            "cid": "18950",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/18950"
        },
        {
            "name": "D-Mannitol",
            "residue": "man+aldi",
            "count": 0,
            "cid": "6251",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6251"
        },
        {
            "name": "N-Acetyl-D-mannosamine",
            "residue": "mannac",
            "count": 0,
            "cid": "439281",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439281"
        },
        {
            "name": "Methyl",
            "residue": "me",
            "count": 0
        },
        {
            "name": "N-Acetyl-Neuraminic Acid",
            "residue": "neuac",
            "count": 2,
            "cid": "439197",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
        },
        {
            "name": "N-Glycolyl-Neuraminic Acid",
            "residue": "neugc",
            "count": 0,
            "cid": "440001",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
        },
        {
            "name": "Phosphate",
            "residue": "p",
            "count": 0,
            "cid": "1061",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
        },
        {
            "name": "Pentose",
            "residue": "pent",
            "count": 0,
            "cid": "229",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
        },
        {
            "name": "Sulfate",
            "residue": "s",
            "count": 0,
            "cid": "1117",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
        },
        {
            "name": "Sialic acid",
            "residue": "sia",
            "count": 2
        },
        {
            "name": "Other",
            "residue": "other",
            "count": 0
        },
        {
            "name": "Floating substituent",
            "residue": "x",
            "count": 0
        },
        {
            "name": "D-Xylose",
            "residue": "xyl",
            "count": 0,
            "cid": "135191",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/135191"
        },
        {
            "name": "Ketodeoxynononic acid",
            "residue": "kdn",
            "count": 0,
            "cid": "13991616",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/13991616"
        }
    ],
    "motifs": [
        {
            "id": "GGM.000001",
            "name": "Type 2 LN",
            "synonym": "N-acetyllactosamine",
            "keywords": "LacNAc; N-acetyllactosamine; N-acetyllactosamine type 2"
        },
        {
            "id": "GGM.000034",
            "name": "3-Sialyl-LN (type 2)",
            "synonym": "",
            "keywords": "LacNAc; N-acetyllactosamine; N-acetyllactosamine type 2"
        },
        {
            "id": "GGM.000056",
            "name": "2,6-Branched O-mannose",
            "synonym": "",
            "keywords": ""
        }
    ],
    "interactions": [],
    "expression": [
        {
            "category": "tissue",
            "tissue": {
                "name": "substantia nigra",
                "namespace": "UBERON",
                "id": "0002038",
                "url": "http://purl.obolibrary.org/obo/UBERON_0002038"
            },
            "cell_line": {},
            "abundance": "",
            "evidence": [
                {
                    "id": "11430",
                    "database": "GlyConnect",
                    "url": "https://glyconnect.expasy.org/browser/structures/11430"
                },
                {
                    "id": "34191522",
                    "database": "PubMed",
                    "url": "https://glygen.org/publication/PubMed/34191522"
                },
                {
                    "id": "40",
                    "database": "GlyConnect",
                    "url": "https://glyconnect.expasy.org/browser/structures/40"
                }
            ]
        }
    ],
    "subsumption": [
        {
            "related_accession": "G05194RK",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G09221QI",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G11461WJ",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G13691FL",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G20253QO",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G23039IV",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G24428CW",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G30945GT",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G39686UR",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G39927PF",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G48275JX",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G63498YL",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G65877AW",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G68729NQ",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G78355JG",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G96496HU",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G05194RK",
            "relationship": "basecomposition"
        },
        {
            "related_accession": "G65877AW",
            "relationship": "composition"
        },
        {
            "related_accession": "G11461WJ",
            "relationship": "subsumedby"
        },
        {
            "related_accession": "G13691FL",
            "relationship": "subsumedby"
        },
        {
            "related_accession": "G63498YL",
            "relationship": "subsumedby"
        },
        {
            "related_accession": "G24428CW",
            "relationship": "topology"
        }
    ],
    "names": [
        {
            "name": "2,6-Branched O-mannose",
            "domain": "Semantic Name"
        },
        {
            "name": "2,6-Branched O-mannose",
            "domain": "motifname"
        }
    ],
    "keywords": [
        "fully_determined"
    ],
    "glycotree_pathways": [
        {
            "source": "G26325MG",
            "target": "G06953DR",
            "residue_affected": {
                "id": "O813",
                "full_name": "α-D-NeupNAc"
            },
            "enzymes": [
                {
                    "uniprotkb_ac": "P54751",
                    "tax_name": "Mus musculus"
                }
            ]
        },
        {
            "source": "G16404NW",
            "target": "G26325MG",
            "residue_affected": {
                "id": "O812",
                "full_name": "α-D-NeupNAc"
            },
            "enzymes": [
                {
                    "uniprotkb_ac": "P54751",
                    "tax_name": "Mus musculus"
                }
            ]
        },
        {
            "source": "G15557BG",
            "target": "G16404NW",
            "residue_affected": {
                "id": "O809",
                "full_name": "β-D-Galp"
            },
            "enzymes": [
                {
                    "uniprotkb_ac": "Q9Z2Y2",
                    "tax_name": "Mus musculus"
                }
            ]
        },
        {
            "source": "G56868BH",
            "target": "G15557BG",
            "residue_affected": {
                "id": "O808",
                "full_name": "β-D-Galp"
            },
            "enzymes": [
                {
                    "uniprotkb_ac": "Q9Z2Y2",
                    "tax_name": "Mus musculus"
                }
            ]
        },
        {
            "source": "G14651SS",
            "target": "G56868BH",
            "residue_affected": {
                "id": "O806",
                "full_name": "β-D-GlcpNAc"
            },
            "enzymes": [
                {
                    "uniprotkb_ac": "Q91X88",
                    "tax_name": "Mus musculus"
                }
            ]
        },
        {
            "source": "G61491DK",
            "target": "G14651SS",
            "residue_affected": {
                "id": "O807",
                "full_name": "β-D-GlcpNAc"
            },
            "enzymes": [
                {
                    "uniprotkb_ac": "Q765H6",
                    "tax_name": "Mus musculus"
                }
            ]
        }
    ],
    "dictionary": {
        "term": "2,6-Branched O-mannose",
        "term_in_sentence": "Collectively, these results strongly suggest that GnT-IX may be a novel beta1,6-N-acetylglucosaminyltransferase that is responsible for the formation of the 2,6-branched structure in the brain O-mannosyl glycan.[PMID:14617637]",
        "publication": "14617637|10469154",
        "definition": "(2S,4S,5R,6R)-5-Acetamido-2-[(2S,3R,4S,5S,6R)-2-[(2R,3S,4R,5R,6R)-5-acetamido-6-[[(2R,3S,4S,5S,6S)-5-[(2S,3R,4R,5S,6R)-3-acetamido-5-[(2S,3R,4S,5S,6R)-4-[(2S,4S,5R,6R)-5-acetamido-2-carboxy-4-hydroxy-6-[(1R,2R)-1,2,3-trihydroxypropyl]oxan-2-yl]oxy-3,5-dihydroxy-6-(hydroxymethyl)oxan-2-yl]oxy-4-hydroxy-6-(hydroxymethyl)oxan-2-yl]oxy-3,4,6-trihydroxyoxan-2-yl]methoxy]-4-hydroxy-2-(hydroxymethyl)oxan-3-yl]oxy-3,5-dihydroxy-6-(hydroxymethyl)oxan-4-yl]oxy-4-hydroxy-6-[(1R,2R)-1,2,3-trihydroxypropyl]oxane-2-carboxylic acid.[CHEBI:157265]",
        "term_xref": "CHEBI:157265|GlycoMotif:GGM.000056|GTC:G06953DR",
        "synonymns": [
            "5-acetamido-3,5-dideoxy-D-glycero-alpha-D-galacto-non-2-ulopyranosylonic acid-(2\u21923)-beta-D-galacto-hexopyranosyl-(1\u21924)-2-acetamido-2-deoxy-beta-D-gluco-hexopyranosyl-(1\u21922)-[5-acetamido-3,5-dideoxy-D-glycero-alpha-D-galacto-non-2-ulopyranosylonic acid-(2\u21923)-beta-D-galacto-hexopyranosyl-(1\u21924)-2-acetamido-2-deoxy-beta-D-gluco-hexopyranosyl-(1\u21926)]-alpha-D-manno-hexopyranose",
            "NeuAc(a2-3)Gal(b1-4)GlcNAc(b1-2)[NeuAc(a2-3)Gal(b1-4)GlcNAc(b1-6)]a-Man"
        ],
        "function": "",
        "disease_associations": "",
        "wikipedia": "",
        "essentials_of_glycobiology": "",
        "evidence": [
            {
                "id": "GSD000001",
                "url": "https://wiki.glygen.org/Glycan_structure_dictionary#GSD000001",
                "database": "Glycan Structure Dictionary"
            }
        ]
    },
    "biomarkers": [],
    "section_stats": [
        {
            "table_id": "glycoprotein",
            "table_stats": [
                {
                    "field": "total",
                    "count": 0
                },
                {
                    "field": "total_sites",
                    "count": 0
                }
            ],
            "sort_fields": []
        },
        {
            "table_id": "expression",
            "table_stats": [
                {
                    "field": "total",
                    "count": 1
                },
                {
                    "field": "total_sites",
                    "count": 0
                }
            ],
            "sort_fields": [
                "category",
                "tissue.name",
                "tissue.namespace",
                "tissue.id",
                "tissue.url",
                "abundance"
            ]
        },
        {
            "table_id": "expression_tissue",
            "table_stats": [
                {
                    "field": "total",
                    "count": 1
                },
                {
                    "field": "total_sites",
                    "count": 0
                }
            ],
            "sort_fields": [
                "category",
                "tissue.name",
                "tissue.namespace",
                "tissue.id",
                "tissue.url",
                "abundance"
            ]
        },
        {
            "table_id": "expression_cell_line",
            "table_stats": [
                {
                    "field": "total",
                    "count": 0
                },
                {
                    "field": "total_sites",
                    "count": 0
                }
            ],
            "sort_fields": [
                "category",
                "abundance"
            ]
        },
        {
            "table_id": "publication",
            "table_stats": [
                {
                    "field": "total",
                    "count": 3
                }
            ],
            "sort_fields": [
                "title",
                "journal",
                "date",
                "authors"
            ]
        }
    ]
}