{
    "glytoucan_ac": "G04576KS",
    "number_monosaccharides": 10,
    "number_monosaccharides_suffix": "",
    "mass": 1890.66,
    "mass_pme": 2367.19,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "yes",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b1-D-Neup5Ac-(2\u21926)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)-\u03b1-D-Manp-(1\u21923)[\u03b1-D-Manp-(1\u21923)[\u03b1-D-Manp-(1\u21926)]-\u03b1-D-Manp-(1\u21926)]-\u03b2-D-Manp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)-\u03b2-D-GlcpNAc-(1\u2192",
    "wurcs": "WURCS=2.0/5,10,9/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O]/1-1-2-3-1-4-5-3-3-3/a4-b1_b4-c1_c3-d1_c6-h1_d2-e1_e4-f1_f6-g2_h3-i1_h6-j1",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 11s:n-acetyl 12b:a-dman-HEX-1:5 13b:a-dman-HEX-1:5 14b:a-dman-HEX-1:5 LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:9o(6+2)10d 10:10d(5+1)11n 11:5o(6+1)12d 12:12o(3+1)13d 13:12o(6+1)14d",
    "inchi": "InChI=1S/C71H118N4O54/c1-16(83)72-31-20(87)5-71(70(109)110,129-57(31)35(89)21(88)6-76)113-15-30-39(93)47(101)51(105)67(122-30)124-55-26(11-81)119-63(34(44(55)98)75-19(4)86)128-60-48(102)38(92)24(9-79)117-69(60)127-59-41(95)29(121-68(53(59)107)125-56-27(12-82)118-62(33(43(56)97)74-18(3)85)123-54-25(10-80)114-61(108)32(42(54)96)73-17(2)84)14-112-65-52(106)58(126-66-50(104)46(100)37(91)23(8-78)116-66)40(94)28(120-65)13-111-64-49(103)45(99)36(90)22(7-77)115-64/h20-69,76-82,87-108H,5-15H2,1-4H3,(H,72,83)(H,73,84)(H,74,85)(H,75,86)(H,109,110)/t20-,21+,22+,23+,24+,25+,26+,27+,28+,29+,30+,31+,32+,33+,34+,35+,36+,37+,38+,39-,40+,41+,42+,43+,44+,45-,46-,47-,48-,49-,50-,51+,52-,53-,54+,55+,56+,57+,58-,59-,60-,61+,62-,63-,64-,65-,66+,67-,68-,69+,71+/m0/s1",
    "glycam": "DNeup5Aca2-6DGalpb1-4DGlcpNAcb1-2DManpa1-3[DManpa1-3[DManpa1-6]DManpa1-6]DManpb1-4DGlcpNAcb1-4DGlcpNAcb1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H](C[C@@](O[C@H]1[C@@H]([C@@H](CO)O)O)(C(=O)O)OC[C@@H]2[C@@H]([C@@H]([C@H]([C@@H](O2)O[C@@H]3[C@H](O[C@H]([C@@H]([C@H]3O)NC(=O)C)O[C@H]4[C@H]([C@@H]([C@H](O[C@@H]4O[C@H]5[C@@H]([C@H](O[C@H]([C@H]5O)O[C@@H]6[C@H](O[C@H]([C@@H]([C@H]6O)NC(=O)C)O[C@@H]7[C@H](O[C@H]([C@@H]([C@H]7O)NC(=O)C)O)CO)CO)CO[C@@H]8[C@H]([C@H]([C@@H]([C@H](O8)CO[C@@H]9[C@H]([C@H]([C@@H]([C@H](O9)CO)O)O)O)O)O[C@@H]1[C@H]([C@H]([C@@H]([C@H](O1)CO)O)O)O)O)O)CO)O)O)CO)O)O)O)O",
    "inchi_key": {
        "key": "QWBBYWKIEBQAQM-MHJJNJBDSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/QWBBYWKIEBQAQM-MHJJNJBDSA-N"
    },
    "byonic": "HexNAc(3)Hex(6)NeuAc(1) % 1890.66104015",
    "gwb": "freeEnd--1b1D-GlcNAc,p--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p--2b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuAc,p)--6a1D-Man,p(--3a1D-Man,p)--6a1D-Man,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "317",
                    "url": "https://glyconnect.expasy.org/browser/structures/317"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G04576KS",
                    "url": "https://glycosmos.org/glycans/show/G04576KS"
                },
                {
                    "database": "PubMed",
                    "id": "33167210",
                    "url": "https://glygen.org/publication/PubMed/33167210"
                },
                {
                    "database": "PubMed",
                    "id": "33383989",
                    "url": "https://glygen.org/publication/PubMed/33383989"
                },
                {
                    "database": "UniCarbKB",
                    "id": "GLY_000341",
                    "url": "http://data.glygen.org/GLY_000341"
                }
            ],
            "common_name": "Human",
            "glygen_name": "Human",
            "reference_species": "Homo sapiens [9606]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Hybrid",
                "url": ""
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15907-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 1",
            "gene": "ST6GAL1",
            "gene_link": "https://www.uniprot.org/uniprot/P15907",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q64685-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 1",
            "gene": "St6gal1",
            "gene_link": "https://www.uniprot.org/uniprot/Q64685",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q76K27-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 2",
            "gene": "St6gal2",
            "gene_link": "https://www.uniprot.org/uniprot/Q76K27",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96JF0-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 2",
            "gene": "ST6GAL2",
            "gene_link": "https://www.uniprot.org/uniprot/Q96JF0",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04576KS"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8K2A8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04576KS",
            "protein_name": "Dol-P-Man:Man(5)GlcNAc(2)-PP-Dol alpha-1,3-mannosyltransferase",
            "gene": "Alg3",
            "gene_link": "https://www.uniprot.org/uniprot/Q8K2A8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04576KS",
                    "database": "SandBox",
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