{
    "glytoucan_ac": "G04483SK",
    "number_monosaccharides": 10,
    "number_monosaccharides_suffix": "",
    "mass": 1786.65,
    "mass_pme": 2221.14,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "yes",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)-\u03b1-D-Manp-(1\u21923)[\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)-\u03b1-D-Manp-(1\u21926)]-\u03b2-D-Manp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)[\u03b1-L-Fucp-(1\u21926)]-\u03b2-D-GlcpNAc-(1\u2192",
    "wurcs": "WURCS=2.0/5,10,9/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][a1221m-1a_1-5]/1-1-2-3-1-4-3-1-4-5/a4-b1_a6-j1_b4-c1_c3-d1_c6-g1_d4-e1_e4-f1_g2-h1_h4-i1",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dman-HEX-1:5 11b:b-dglc-HEX-1:5 12s:n-acetyl 13b:b-dgal-HEX-1:5 14b:a-lgal-HEX-1:5|6:d LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(4+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:5o(6+1)10d 10:10o(2+1)11d 11:11d(2+1)12n 12:11o(4+1)13d 13:1o(6+1)14d",
    "inchi": "InChI=1S/C68H114N4O50/c1-15-33(84)42(93)47(98)63(106-15)104-14-28-56(38(89)29(59(103)107-28)69-16(2)80)117-61-31(71-18(4)82)40(91)54(25(11-78)112-61)120-67-51(102)57(121-66-50(101)46(97)55(26(12-79)114-66)116-60-30(70-17(3)81)39(90)52(23(9-76)111-60)118-64-48(99)43(94)34(85)20(6-73)108-64)37(88)27(115-67)13-105-68-58(45(96)36(87)22(8-75)110-68)122-62-32(72-19(5)83)41(92)53(24(10-77)113-62)119-65-49(100)44(95)35(86)21(7-74)109-65/h15,20-68,73-79,84-103H,6-14H2,1-5H3,(H,69,80)(H,70,81)(H,71,82)(H,72,83)/t15-,20+,21+,22+,23+,24+,25+,26+,27+,28+,29+,30+,31+,32+,33+,34-,35-,36+,37+,38+,39+,40+,41+,42+,43-,44-,45-,46+,47-,48+,49+,50-,51-,52+,53+,54+,55+,56+,57-,58-,59+,60-,61-,62-,63+,64-,65-,66+,67-,68-/m0/s1",
    "glycam": "DGalpb1-4DGlcpNAcb1-4DManpa1-3[DGalpb1-4DGlcpNAcb1-2DManpa1-6]DManpb1-4DGlcpNAcb1-4[LFucpa1-6]DGlcpNAcb1-OH",
    "smiles_isomeric": "C[C@H]1[C@H]([C@H]([C@@H]([C@@H](O1)OC[C@@H]2[C@H]([C@@H]([C@H]([C@@H](O2)O)NC(=O)C)O)O[C@H]3[C@@H]([C@H]([C@@H]([C@H](O3)CO)O[C@H]4[C@H]([C@H]([C@@H]([C@H](O4)CO[C@@H]5[C@H]([C@H]([C@@H]([C@H](O5)CO)O)O)O[C@H]6[C@@H]([C@H]([C@@H]([C@H](O6)CO)O[C@H]7[C@@H]([C@H]([C@H]([C@H](O7)CO)O)O)O)O)NC(=O)C)O)O[C@@H]8[C@H]([C@H]([C@@H]([C@H](O8)CO)O[C@H]9[C@@H]([C@H]([C@@H]([C@H](O9)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O)O)O)NC(=O)C)O)O)O)O)NC(=O)C)O)O)O",
    "inchi_key": {
        "key": "GACPKDGFLORSAU-JTQLCSPLSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/GACPKDGFLORSAU-JTQLCSPLSA-N"
    },
    "byonic": "HexNAc(4)Hex(5)dHex(1) % 1786.65008149",
    "gwb": "freeEnd--1b1D-GlcNAc,p(--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p--4b1D-GlcNAc,p--4b1D-Gal,p)--6a1D-Man,p--2b1D-GlcNAc,p--4b1D-Gal,p)--6a1L-Fuc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "2005",
                    "url": "https://glyconnect.expasy.org/browser/structures/2005"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G04483SK",
                    "url": "https://glycosmos.org/glycans/show/G04483SK"
                },
                {
                    "database": "PubMed",
                    "id": "11180632",
                    "url": "https://glygen.org/publication/PubMed/11180632"
                },
                {
                    "database": "PubMed",
                    "id": "2110822",
                    "url": "https://glygen.org/publication/PubMed/2110822"
                },
                {
                    "database": "PubMed",
                    "id": "8344278",
                    "url": "https://glygen.org/publication/PubMed/8344278"
                },
                {
                    "database": "UniCarbKB",
                    "id": "GLY_000341",
                    "url": "http://data.glygen.org/GLY_000341"
                }
            ],
            "common_name": "Human",
            "glygen_name": "Human",
            "reference_species": "Homo sapiens [9606]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Biantennary"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Complex",
                "url": ""
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Core-fucosylated"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10469-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q10469",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921V5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat2",
            "gene_link": "https://www.uniprot.org/uniprot/Q921V5",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q812F8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase B",
            "gene": "Mgat4b",
            "gene_link": "https://www.uniprot.org/uniprot/Q812F8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q812G0-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase A",
            "gene": "Mgat4a",
            "gene_link": "https://www.uniprot.org/uniprot/Q812G0",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D306-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase C",
            "gene": "Mgat4c",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D306",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9UBM8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04483SK",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase C",
            "gene": "MGAT4C",
            "gene_link": "https://www.uniprot.org/uniprot/Q9UBM8",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04483SK",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04483SK"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9UM21-1",
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