{
    "glytoucan_ac": "G03542LT",
    "number_monosaccharides": 15,
    "number_monosaccharides_suffix": "",
    "mass": 3170.11,
    "mass_pme": 3940.97,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "no",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "",
    "missing_score": 65,
    "glycan_type": "Saccharide",
    "iupac": "\u03b1-D-Neup5Ac-(2\u21923)[\u03b1-D-Neup5Ac-(2\u21923)-\u03b2-D-Galp-(1\u21924)]-\u03b2-D-GlcpNAc-(1\u21922)[\u03b1-D-Neup5Ac-(2\u21923)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)]-\u03b1-D-Manp-(1\u21923)[\u03b1-D-Neup5Ac-(2\u21923)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)-\u03b1-D-Manp-(1\u21926)]-\u03b2-D-Manp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)-?-D-GlcpNAc-(1\u2192",
    "wurcs": "WURCS=2.0/6,15,14/[a2122h-1x_1-5_2*NCC/3=O][a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][Aad21122h-2a_2-6_5*NCC/3=O][a2112h-1b_1-5]/1-2-3-4-2-5-6-5-2-6-5-4-2-6-5/a4-b1_b4-c1_c3-d1_c6-l1_d2-e1_d4-i1_e3-f2_e4-g1_g3-h2_i4-j1_j3-k2_l2-m1_m4-n1_n3-o2",
    "glycoct": "RES 1b:x-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 10s:n-acetyl 11b:b-dgal-HEX-1:5 12b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 13s:n-acetyl 14b:b-dglc-HEX-1:5 15s:n-acetyl 16b:b-dgal-HEX-1:5 17b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 18s:n-acetyl 19b:a-dman-HEX-1:5 20b:b-dglc-HEX-1:5 21s:n-acetyl 22b:b-dgal-HEX-1:5 23b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 24s:n-acetyl LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(3+2)9d 9:9d(5+1)10n 10:7o(4+1)11d 11:11o(3+2)12d 12:12d(5+1)13n 13:6o(4+1)14d 14:14d(2+1)15n 15:14o(4+1)16d 16:16o(3+2)17d 17:17d(5+1)18n 18:5o(6+1)19d 19:19o(2+1)20d 20:20d(2+1)21n 21:20o(4+1)22d 22:22o(3+2)23d 23:23d(5+1)24n",
    "inchi": "InChI=1S/C120H195N9O88/c1-29(144)121-57-38(153)10-117(113(181)182,210-91(57)66(161)42(157)14-130)214-95-65(129-37(9)152)106(198-55(27-143)90(95)206-110-84(179)99(73(168)49(21-137)193-110)217-120(116(187)188)13-41(156)60(124-32(4)147)94(213-120)69(164)45(160)17-133)209-101-80(175)89(202-104-63(127-35(7)150)77(172)87(52(24-140)196-104)204-108-82(177)97(71(166)47(19-135)191-108)215-118(114(183)184)11-39(154)58(122-30(2)145)92(211-118)67(162)43(158)15-131)54(26-142)199-112(101)207-96-74(169)56(200-107(81(96)176)203-86-51(23-139)195-103(62(76(86)171)126-34(6)149)201-85-50(22-138)190-102(180)61(75(85)170)125-33(5)148)28-189-111-100(79(174)70(165)46(18-134)194-111)208-105-64(128-36(8)151)78(173)88(53(25-141)197-105)205-109-83(178)98(72(167)48(20-136)192-109)216-119(115(185)186)12-40(155)59(123-31(3)146)93(212-119)68(163)44(159)16-132/h38-112,130-143,153-180H,10-28H2,1-9H3,(H,121,144)(H,122,145)(H,123,146)(H,124,147)(H,125,148)(H,126,149)(H,127,150)(H,128,151)(H,129,152)(H,181,182)(H,183,184)(H,185,186)(H,187,188)/t38-,39-,40-,41-,42+,43+,44+,45+,46+,47+,48+,49+,50+,51+,52+,53+,54+,55+,56+,57+,58+,59+,60+,61+,62+,63+,64+,65+,66+,67+,68+,69+,70+,71-,72-,73-,74+,75+,76+,77+,78+,79-,80-,81-,82+,83+,84+,85+,86+,87+,88+,89+,90+,91+,92+,93+,94+,95+,96-,97-,98-,99-,100-,101-,102?,103-,104-,105-,106-,107-,108-,109-,110-,111-,112+,117-,118-,119-,120-/m0/s1",
    "glycam": "",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H](C[C@@](O[C@H]1[C@@H]([C@@H](CO)O)O)(C(=O)O)O[C@@H]2[C@H]([C@@H](O[C@@H]([C@H]2O[C@H]3[C@@H]([C@H]([C@H]([C@H](O3)CO)O)O[C@@]4(C[C@@H]([C@H]([C@@H](O4)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)CO)O[C@H]5[C@H]([C@@H]([C@H](O[C@@H]5O[C@H]6[C@@H]([C@H](O[C@H]([C@H]6O)O[C@@H]7[C@H](O[C@H]([C@@H]([C@H]7O)NC(=O)C)O[C@@H]8[C@H](OC([C@@H]([C@H]8O)NC(=O)C)O)CO)CO)CO[C@@H]9[C@H]([C@H]([C@@H]([C@H](O9)CO)O)O)O[C@H]1[C@@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O[C@@]1(C[C@@H]([C@H]([C@@H](O1)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)NC(=O)C)O)CO)O[C@H]1[C@@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O[C@@]1(C[C@@H]([C@H]([C@@H](O1)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)NC(=O)C)O)NC(=O)C)O",
    "inchi_key": {
        "key": "YHNRICRJBKTZFF-XUEIZHKHSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/YHNRICRJBKTZFF-XUEIZHKHSA-N"
    },
    "byonic": "HexNAc(5)Hex(6)NeuAc(4) % 3170.10603527",
    "gwb": "freeEnd--1?1D-GlcNAc,p--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p(--2b1D-GlcNAc,p(--3a2D-NeuAc,p)--4b1D-Gal,p--3a2D-NeuAc,p)--4b1D-GlcNAc,p--4b1D-Gal,p--3a2D-NeuAc,p)--6a1D-Man,p--2b1D-GlcNAc,p--4b1D-Gal,p--3a2D-NeuAc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9913,
            "name": "Bos taurus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G03542LT",
                    "url": "https://glycosmos.org/glycans/show/G03542LT"
                }
            ],
            "common_name": "Bovine",
            "glygen_name": "Bovine",
            "reference_species": "Bos taurus [9913]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Complex",
                "url": ""
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Triantennary"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10469-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q10469",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03542LT",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03542LT"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921V5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03542LT",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat2",
            "gene_link": "https://www.uniprot.org/uniprot/Q921V5",
            "tax_id": 10090,
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