{
    "glytoucan_ac": "G02106OV",
    "number_monosaccharides": 18,
    "number_monosaccharides_suffix": "",
    "mass": 3294.17,
    "mass_pme": 4093.06,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "no",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "no",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b1-D-Neup5Ac-(2\u21926)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)[\u03b1-D-Galp-(1\u21923)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)]-\u03b1-D-Manp-(1\u21923)[\u03b1-D-Galp-(1\u21923)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)[\u03b1-D-Galp-(1\u21923)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21926)]-\u03b1-D-Manp-(1\u21926)]-\u03b2-D-Manp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)[\u03b1-L-Fucp-(1\u21926)]-\u03b2-D-GlcpNAc-(1\u2192",
    "wurcs": "WURCS=2.0/7,18,17/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O][a2112h-1a_1-5][a1221m-1a_1-5]/1-1-2-3-1-4-5-1-4-6-3-1-4-6-1-4-6-7/a4-b1_a6-r1_b4-c1_c3-d1_c6-k1_d2-e1_d4-h1_e4-f1_f6-g2_h4-i1_i3-j1_k2-l1_k6-o1_l4-m1_m3-n1_o4-p1_p3-q1",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 11s:n-acetyl 12b:b-dglc-HEX-1:5 13s:n-acetyl 14b:b-dgal-HEX-1:5 15b:a-dgal-HEX-1:5 16b:a-dman-HEX-1:5 17b:b-dglc-HEX-1:5 18s:n-acetyl 19b:b-dgal-HEX-1:5 20b:a-dgal-HEX-1:5 21b:b-dglc-HEX-1:5 22s:n-acetyl 23b:b-dgal-HEX-1:5 24b:a-dgal-HEX-1:5 25b:a-lgal-HEX-1:5|6:d LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:9o(6+2)10d 10:10d(5+1)11n 11:6o(4+1)12d 12:12d(2+1)13n 13:12o(4+1)14d 14:14o(3+1)15d 15:5o(6+1)16d 16:16o(2+1)17d 17:17d(2+1)18n 18:17o(4+1)19d 19:19o(3+1)20d 20:16o(6+1)21d 21:21d(2+1)22n 22:21o(4+1)23d 23:23o(3+1)24d 24:1o(6+1)25d",
    "inchi": "InChI=1S/C125H207N7O93/c1-27-60(155)77(172)83(178)113(195-27)192-25-52-99(71(166)54(107(188)196-52)127-29(3)147)213-110-57(130-32(6)150)74(169)97(46(20-143)205-110)218-121-92(187)104(70(165)50(209-121)24-193-122-105(223-111-58(131-33(7)151)76(171)96(47(21-144)206-111)217-120-91(186)103(69(164)42(16-139)202-120)221-116-86(181)80(175)64(159)39(13-136)199-116)82(177)66(161)49(211-122)23-191-108-55(128-30(4)148)72(167)93(43(17-140)203-108)215-118-89(184)101(67(162)40(14-137)200-118)219-114-84(179)78(173)62(157)37(11-134)197-114)222-123-106(88(183)98(48(22-145)208-123)212-109-56(129-31(5)149)73(168)95(45(19-142)204-109)216-119-90(185)102(68(163)41(15-138)201-119)220-115-85(180)79(174)63(158)38(12-135)198-115)224-112-59(132-34(8)152)75(170)94(44(18-141)207-112)214-117-87(182)81(176)65(160)51(210-117)26-194-125(124(189)190)9-35(153)53(126-28(2)146)100(225-125)61(156)36(154)10-133/h27,35-123,133-145,153-188H,9-26H2,1-8H3,(H,126,146)(H,127,147)(H,128,148)(H,129,149)(H,130,150)(H,131,151)(H,132,152)(H,189,190)/t27-,35-,36+,37+,38+,39+,40+,41+,42+,43+,44+,45+,46+,47+,48+,49+,50+,51+,52+,53+,54+,55+,56+,57+,58+,59+,60+,61+,62-,63-,64-,65-,66+,67-,68-,69-,70+,71+,72+,73+,74+,75+,76+,77+,78-,79-,80-,81-,82-,83-,84+,85+,86+,87+,88-,89+,90+,91+,92-,93+,94+,95+,96+,97+,98+,99+,100+,101-,102-,103-,104-,105-,106-,107+,108+,109-,110-,111-,112-,113+,114+,115+,116+,117-,118-,119-,120-,121-,122-,123+,125+/m0/s1",
    "glycam": "DNeup5Aca2-6DGalpb1-4DGlcpNAcb1-2[DGalpa1-3DGalpb1-4DGlcpNAcb1-4]DManpa1-3[DGalpa1-3DGalpb1-4DGlcpNAcb1-2[DGalpa1-3DGalpb1-4DGlcpNAcb1-6]DManpa1-6]DManpb1-4DGlcpNAcb1-4[LFucpa1-6]DGlcpNAcb1-OH",
    "smiles_isomeric": "C[C@H]1[C@H]([C@H]([C@@H]([C@@H](O1)OC[C@@H]2[C@H]([C@@H]([C@H]([C@@H](O2)O)NC(=O)C)O)O[C@H]3[C@@H]([C@H]([C@@H]([C@H](O3)CO)O[C@H]4[C@H]([C@H]([C@@H]([C@H](O4)CO[C@@H]5[C@H]([C@H]([C@@H]([C@H](O5)CO[C@H]6[C@@H]([C@H]([C@@H]([C@H](O6)CO)O[C@H]7[C@@H]([C@H]([C@H]([C@H](O7)CO)O)O[C@@H]8[C@@H]([C@H]([C@H]([C@H](O8)CO)O)O)O)O)O)NC(=O)C)O)O)O[C@H]9[C@@H]([C@H]([C@@H]([C@H](O9)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O[C@@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O)O)O)O)NC(=O)C)O)O[C@@H]1[C@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O[C@@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O)O)O)O)NC(=O)C)O)O[C@H]1[C@@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO[C@@]1(C[C@@H]([C@H]([C@@H](O1)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)O)O)NC(=O)C)O)O)NC(=O)C)O)O)O",
    "inchi_key": {
        "key": "ZFPFQJFNJGPEAZ-CVTQSPOTSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/ZFPFQJFNJGPEAZ-CVTQSPOTSA-N"
    },
    "byonic": "HexNAc(6)Hex(10)dHex(1)NeuAc(1) % 3294.16836083",
    "gwb": "freeEnd--1b1D-GlcNAc,p(--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p(--2b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuAc,p)--4b1D-GlcNAc,p--4b1D-Gal,p--3a1D-Gal,p)--6a1D-Man,p(--2b1D-GlcNAc,p--4b1D-Gal,p--3a1D-Gal,p)--6b1D-GlcNAc,p--4b1D-Gal,p--3a1D-Gal,p)--6a1L-Fuc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9913,
            "name": "Bos taurus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G02106OV",
                    "url": "https://glycosmos.org/glycans/show/G02106OV"
                }
            ],
            "common_name": "Bovine",
            "glygen_name": "Bovine",
            "reference_species": "Bos taurus [9913]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Complex",
                "url": ""
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Core-fucosylated"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Tetraantennary"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G02106OV",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G02106OV",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G02106OV"
                }
            ]
        },
        {
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