{
    "glytoucan_ac": "G00666EC",
    "number_monosaccharides": 18,
    "number_monosaccharides_suffix": "",
    "mass": 3520.26,
    "mass_pme": 4347.19,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "yes",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b1-D-Neup5Ac-(2\u21926)[\u03b1-L-Fucp-(1\u21922)]-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)[\u03b1-D-Neup5Ac-(2\u21926)[\u03b1-L-Fucp-(1\u21922)]-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21926)]-\u03b1-D-Manp-(1\u21923)[\u03b2-D-GlcpNAc-(1\u21924)][\u03b1-D-Neup5Ac-(2\u21926)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)-\u03b1-D-Manp-(1\u21926)]-\u03b2-D-Manp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)[\u03b1-L-Fucp-(1\u21926)]-\u03b2-D-GlcpNAc-(1\u2192",
    "wurcs": "WURCS=2.0/6,18,17/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][a1221m-1a_1-5][Aad21122h-2a_2-6_5*NCC/3=O]/1-1-2-3-1-4-5-6-1-4-5-6-1-3-1-4-6-5/a4-b1_a6-r1_b4-c1_c3-d1_c4-m1_c6-n1_d2-e1_d6-i1_e4-f1_f2-g1_f6-h2_i4-j1_j2-k1_j6-l2_n2-o1_o4-p1_p6-q2",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-lgal-HEX-1:5|6:d 11b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 12s:n-acetyl 13b:b-dglc-HEX-1:5 14s:n-acetyl 15b:b-dgal-HEX-1:5 16b:a-lgal-HEX-1:5|6:d 17b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 18s:n-acetyl 19b:b-dglc-HEX-1:5 20s:n-acetyl 21b:a-dman-HEX-1:5 22b:b-dglc-HEX-1:5 23s:n-acetyl 24b:b-dgal-HEX-1:5 25b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 26s:n-acetyl 27b:a-lgal-HEX-1:5|6:d LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:9o(2+1)10d 10:9o(6+2)11d 11:11d(5+1)12n 12:6o(6+1)13d 13:13d(2+1)14n 14:13o(4+1)15d 15:15o(2+1)16d 16:15o(6+2)17d 17:17d(5+1)18n 18:5o(4+1)19d 19:19d(2+1)20n 20:5o(6+1)21d 21:21o(2+1)22d 22:22d(2+1)23n 23:22o(4+1)24d 24:24o(6+2)25d 25:25d(5+1)26n 26:1o(6+1)27d",
    "inchi": "InChI=1S/C135H221N9O97/c1-31-70(169)88(187)96(195)121(213-31)208-26-59-105(83(182)64(115(200)216-59)139-37(7)157)228-118-67(142-40(10)160)85(184)102(53(23-152)220-118)230-125-100(199)110(106(233-117-65(140-38(8)158)82(181)76(175)49(19-148)217-117)60(227-125)27-209-126-111(92(191)77(176)50(20-149)218-126)235-119-68(143-41(11)161)86(185)101(52(22-151)221-119)229-124-99(198)91(190)78(177)56(223-124)28-210-133(130(201)202)13-43(163)61(136-34(4)154)107(239-133)73(172)46(166)16-145)234-129-112(236-120-69(144-42(12)162)87(186)104(54(24-153)222-120)232-128-114(238-123-98(197)90(189)72(171)33(3)215-123)95(194)81(180)58(226-128)30-212-135(132(205)206)15-45(165)63(138-36(6)156)109(241-135)75(174)48(168)18-147)93(192)79(178)55(224-129)25-207-116-66(141-39(9)159)84(183)103(51(21-150)219-116)231-127-113(237-122-97(196)89(188)71(170)32(2)214-122)94(193)80(179)57(225-127)29-211-134(131(203)204)14-44(164)62(137-35(5)155)108(240-134)74(173)47(167)17-146/h31-33,43-129,145-153,163-200H,13-30H2,1-12H3,(H,136,154)(H,137,155)(H,138,156)(H,139,157)(H,140,158)(H,141,159)(H,142,160)(H,143,161)(H,144,162)(H,201,202)(H,203,204)(H,205,206)/t31-,32-,33-,43-,44-,45-,46+,47+,48+,49+,50+,51+,52+,53+,54+,55+,56+,57+,58+,59+,60+,61+,62+,63+,64+,65+,66+,67+,68+,69+,70+,71+,72+,73+,74+,75+,76+,77+,78-,79+,80-,81-,82+,83+,84+,85+,86+,87+,88+,89+,90+,91-,92-,93-,94-,95-,96-,97-,98-,99+,100-,101+,102+,103+,104+,105+,106+,107+,108+,109+,110+,111-,112-,113+,114+,115+,116+,117-,118-,119-,120-,121+,122-,123-,124-,125-,126-,127-,128-,129+,133+,134+,135+/m0/s1",
    "glycam": "DNeup5Aca2-6[LFucpa1-2]DGalpb1-4DGlcpNAcb1-2[DNeup5Aca2-6[LFucpa1-2]DGalpb1-4DGlcpNAcb1-6]DManpa1-3[DNeup5Aca2-6DGalpb1-4DGlcpNAcb1-2DManpa1-6][DGlcpNAcb1-4]DManpb1-4DGlcpNAcb1-4[LFucpa1-6]DGlcpNAcb1-OH",
    "smiles_isomeric": "C[C@H]1[C@H]([C@H]([C@@H]([C@@H](O1)OC[C@@H]2[C@H]([C@@H]([C@H]([C@@H](O2)O)NC(=O)C)O)O[C@H]3[C@@H]([C@H]([C@@H]([C@H](O3)CO)O[C@H]4[C@H]([C@H]([C@@H]([C@H](O4)CO[C@@H]5[C@H]([C@H]([C@@H]([C@H](O5)CO)O)O)O[C@H]6[C@@H]([C@H]([C@@H]([C@H](O6)CO)O[C@H]7[C@@H]([C@H]([C@H]([C@H](O7)CO[C@@]8(C[C@@H]([C@H]([C@@H](O8)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)O)O)NC(=O)C)O[C@H]9[C@@H]([C@H]([C@@H]([C@H](O9)CO)O)O)NC(=O)C)O[C@@H]1[C@H]([C@H]([C@@H]([C@H](O1)CO[C@H]1[C@@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO[C@@]1(C[C@@H]([C@H]([C@@H](O1)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)O[C@H]1[C@H]([C@@H]([C@@H]([C@@H](O1)C)O)O)O)O)NC(=O)C)O)O)O[C@H]1[C@@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO[C@@]1(C[C@@H]([C@H]([C@@H](O1)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)O[C@H]1[C@H]([C@@H]([C@@H]([C@@H](O1)C)O)O)O)O)NC(=O)C)O)O)NC(=O)C)O)O)O",
    "inchi_key": {
        "key": "LRDNMLMXAVWBCE-AZBIDXEMSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/LRDNMLMXAVWBCE-AZBIDXEMSA-N"
    },
    "byonic": "HexNAc(6)Hex(6)dHex(3)NeuAc(3) % 3520.26371784",
    "gwb": "freeEnd--1b1D-GlcNAc,p(--4b1D-GlcNAc,p--4b1D-Man,p((--3a1D-Man,p(--2b1D-GlcNAc,p--4b1D-Gal,p(--2a1L-Fuc,p)--6a2D-NeuAc,p)--6b1D-GlcNAc,p--4b1D-Gal,p(--2a1L-Fuc,p)--6a2D-NeuAc,p)--4b1D-GlcNAc,p)--6a1D-Man,p--2b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuAc,p)--6a1L-Fuc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "2402",
                    "url": "https://glyconnect.expasy.org/browser/structures/2402"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G00666EC",
                    "url": "https://glycosmos.org/glycans/show/G00666EC"
                },
                {
                    "database": "PubMed",
                    "id": "7763244",
                    "url": "https://glygen.org/publication/PubMed/7763244"
                }
            ],
            "common_name": "Human",
            "glygen_name": "Human",
            "reference_species": "Homo sapiens [9606]"
        },
        {
            "taxid": 10114,
            "name": "Rattus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G00666EC",
                    "url": "https://glycosmos.org/glycans/show/G00666EC"
                }
            ],
            "common_name": "Rattus",
            "glygen_name": "Rattus",
            "reference_species": "Rattus [10114]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Bisected"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Complex",
                "url": ""
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Core-fucosylated"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Triantennary"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00666EC",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00666EC"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00666EC",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
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