{
    "glytoucan_ac": "G00033MO",
    "number_monosaccharides": 3,
    "number_monosaccharides_suffix": "",
    "mass": 586.22,
    "mass_pme": 740.39,
    "tool_support": {
        "gnome_glygen_oglycans": "yes",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "yes",
        "pdb": "yes",
        "gnome_glygen_nglycans": "no",
        "gnome_glycotree_nglycans": "no"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b2-D-Galp-(1\u21923)[\u03b2-D-GlcpNAc-(1\u21926)]-\u03b1-D-GalpNAc-(1\u2192",
    "wurcs": "WURCS=2.0/3,3,2/[a2112h-1a_1-5_2*NCC/3=O][a2112h-1b_1-5][a2122h-1b_1-5_2*NCC/3=O]/1-2-3/a3-b1_a6-c1",
    "glycoct": "RES 1b:a-dgal-HEX-1:5 2s:n-acetyl 3b:b-dgal-HEX-1:5 4b:b-dglc-HEX-1:5 5s:n-acetyl LIN 1:1d(2+1)2n 2:1o(3+1)3d 3:1o(6+1)4d 4:4d(2+1)5n",
    "inchi": "InChI=1S/C22H38N2O16/c1-6(27)23-11-16(32)13(29)8(3-25)38-21(11)36-5-10-15(31)19(12(20(35)37-10)24-7(2)28)40-22-18(34)17(33)14(30)9(4-26)39-22/h8-22,25-26,29-35H,3-5H2,1-2H3,(H,23,27)(H,24,28)/t8-,9-,10-,11-,12-,13-,14+,15+,16-,17+,18-,19-,20+,21-,22+/m1/s1",
    "glycam": "DGalpb1-3[DGlcpNAcb1-6]DGalpNAca1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H]([C@@H]([C@H](O[C@H]1OC[C@@H]2[C@@H]([C@@H]([C@H]([C@H](O2)O)NC(=O)C)O[C@H]3[C@@H]([C@H]([C@H]([C@H](O3)CO)O)O)O)O)CO)O)O",
    "inchi_key": {
        "key": "AULDNGHDBFNGGC-XYVIJKTMSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/AULDNGHDBFNGGC-XYVIJKTMSA-N"
    },
    "byonic": "HexNAc(2)Hex(1) % 586.22213341",
    "gwb": "freeEnd--1a1D-GalNAc,p(--3b1D-Gal,p)--6b1D-GlcNAc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9913,
            "name": "Bos taurus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3318",
                    "url": "https://glyconnect.expasy.org/browser/structures/3318"
                },
                {
                    "database": "PubMed",
                    "id": "1718265",
                    "url": "https://glygen.org/publication/PubMed/1718265"
                },
                {
                    "database": "PubMed",
                    "id": "1730232",
                    "url": "https://glygen.org/publication/PubMed/1730232"
                },
                {
                    "database": "PubMed",
                    "id": "3768900",
                    "url": "https://glygen.org/publication/PubMed/3768900"
                },
                {
                    "database": "PubMed",
                    "id": "9874229",
                    "url": "https://glygen.org/publication/PubMed/9874229"
                }
            ],
            "common_name": "Bovine",
            "glygen_name": "Bovine",
            "reference_species": "Bos taurus [9913]"
        },
        {
            "taxid": 7227,
            "name": "Drosophila melanogaster",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3318",
                    "url": "https://glyconnect.expasy.org/browser/structures/3318"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G00033MO",
                    "url": "https://glycosmos.org/glycans/show/G00033MO"
                },
                {
                    "database": "PubMed",
                    "id": "18725413",
                    "url": "https://glygen.org/publication/PubMed/18725413"
                }
            ],
            "common_name": "Fruit fly",
            "glygen_name": "Fruit fly",
            "reference_species": "Drosophila melanogaster [7227]"
        },
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3318",
                    "url": "https://glyconnect.expasy.org/browser/structures/3318"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G00033MO",
                    "url": "https://glycosmos.org/glycans/show/G00033MO"
                },
                {
                    "database": "PubMed",
                    "id": "14673092",
                    "url": "https://glygen.org/publication/PubMed/14673092"
                },
                {
                    "database": "PubMed",
                    "id": "1488058",
                    "url": "https://glygen.org/publication/PubMed/1488058"
                },
                {
                    "database": "PubMed",
                    "id": "1627559",
                    "url": "https://glygen.org/publication/PubMed/1627559"
                },
                {
                    "database": "PubMed",
                    "id": "25187573",
                    "url": "https://glygen.org/publication/PubMed/25187573"
                },
                {
                    "database": "PubMed",
                    "id": "2580706",
                    "url": "https://glygen.org/publication/PubMed/2580706"
                },
                {
                    "database": "PubMed",
                    "id": "3003076",
                    "url": "https://glygen.org/publication/PubMed/3003076"
                },
                {
                    "database": "PubMed",
                    "id": "32310329",
                    "url": "https://glygen.org/publication/PubMed/32310329"
                },
                {
                    "database": "PubMed",
                    "id": "32841605",
                    "url": "https://glygen.org/publication/PubMed/32841605"
                },
                {
                    "database": "PubMed",
                    "id": "3345749",
                    "url": "https://glygen.org/publication/PubMed/3345749"
                },
                {
                    "database": "PubMed",
                    "id": "34191522",
                    "url": "https://glygen.org/publication/PubMed/34191522"
                },
                {
                    "database": "PubMed",
                    "id": "3956483",
                    "url": "https://glygen.org/publication/PubMed/3956483"
                },
                {
                    "database": "PubMed",
                    "id": "4357339",
                    "url": "https://glygen.org/publication/PubMed/4357339"
                },
                {
                    "database": "PubMed",
                    "id": "6539332",
                    "url": "https://glygen.org/publication/PubMed/6539332"
                },
                {
                    "database": "PubMed",
                    "id": "7080930",
                    "url": "https://glygen.org/publication/PubMed/7080930"
                },
                {
                    "database": "PubMed",
                    "id": "7140761",
                    "url": "https://glygen.org/publication/PubMed/7140761"
                },
                {
                    "database": "PubMed",
                    "id": "7215351",
                    "url": "https://glygen.org/publication/PubMed/7215351"
                },
                {
                    "database": "PubMed",
                    "id": "952526",
                    "url": "https://glygen.org/publication/PubMed/952526"
                },
                {
                    "database": "PubMed",
                    "id": "9530955",
                    "url": "https://glygen.org/publication/PubMed/9530955"
                }
            ],
            "common_name": "Human",
            "glygen_name": "Human",
            "reference_species": "Homo sapiens [9606]"
        },
        {
            "taxid": 10090,
            "name": "Mus musculus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3318",
                    "url": "https://glyconnect.expasy.org/browser/structures/3318"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G00033MO",
                    "url": "https://glycosmos.org/glycans/show/G00033MO"
                },
                {
                    "database": "PubMed",
                    "id": "14673092",
                    "url": "https://glygen.org/publication/PubMed/14673092"
                },
                {
                    "database": "PubMed",
                    "id": "35022400",
                    "url": "https://glygen.org/publication/PubMed/35022400"
                }
            ],
            "common_name": "Mouse",
            "glygen_name": "Mouse",
            "reference_species": "Mus musculus [10090]"
        },
        {
            "taxid": 9823,
            "name": "Sus scrofa",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3318",
                    "url": "https://glyconnect.expasy.org/browser/structures/3318"
                },
                {
                    "database": "PubMed",
                    "id": "6436240",
                    "url": "https://glygen.org/publication/PubMed/6436240"
                }
            ],
            "common_name": "Pig",
            "glygen_name": "Pig",
            "reference_species": "Sus scrofa [9823]"
        },
        {
            "taxid": 10116,
            "name": "Rattus norvegicus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3318",
                    "url": "https://glyconnect.expasy.org/browser/structures/3318"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G00033MO",
                    "url": "https://glycosmos.org/glycans/show/G00033MO"
                },
                {
                    "database": "PubMed",
                    "id": "6882773",
                    "url": "https://glygen.org/publication/PubMed/6882773"
                },
                {
                    "database": "PubMed",
                    "id": "7061510",
                    "url": "https://glygen.org/publication/PubMed/7061510"
                }
            ],
            "common_name": "Rat",
            "glygen_name": "Rat",
            "reference_species": "Rattus norvegicus [10116]"
        },
        {
            "taxid": 2697049,
            "name": "Severe acute respiratory syndrome coronavirus 2",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3318",
                    "url": "https://glyconnect.expasy.org/browser/structures/3318"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G00033MO",
                    "url": "https://glycosmos.org/glycans/show/G00033MO"
                },
                {
                    "database": "PubMed",
                    "id": "33871970",
                    "url": "https://glygen.org/publication/PubMed/33871970"
                }
            ],
            "common_name": "2019-nCoV",
            "glygen_name": "SARS-CoV-2",
            "reference_species": "Severe acute respiratory syndrome coronavirus 2 [2697049]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "O-linked",
                "url": ""
            },
            "subtype": {
                "name": "Core 2",
                "url": ""
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "E5D8G1-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase",
            "gene": "Galntl6",
            "gene_link": "https://www.uniprot.org/uniprot/E5D8G1",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "G3X942-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase",
            "gene": "Galnt9",
            "gene_link": "https://www.uniprot.org/uniprot/G3X942",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
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                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O08832-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 4",
            "gene": "Galnt4",
            "gene_link": "https://www.uniprot.org/uniprot/O08832",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O08912-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 1",
            "gene": "Galnt1",
            "gene_link": "https://www.uniprot.org/uniprot/O08912",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P70419-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 3",
            "gene": "Galnt3",
            "gene_link": "https://www.uniprot.org/uniprot/P70419",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
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            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10471-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 2",
            "gene": "GALNT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q10471",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10472-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 1",
            "gene": "GALNT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q10472",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q14435-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 3",
            "gene": "GALNT3",
            "gene_link": "https://www.uniprot.org/uniprot/Q14435",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
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                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q49A17-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase-like 6",
            "gene": "GALNTL6",
            "gene_link": "https://www.uniprot.org/uniprot/Q49A17",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
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                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
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            ]
        },
        {
            "uniprot_canonical_ac": "Q6IS24-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 17",
            "gene": "GALNT17",
            "gene_link": "https://www.uniprot.org/uniprot/Q6IS24",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
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                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
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        },
        {
            "uniprot_canonical_ac": "Q6P9A2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 18",
            "gene": "GALNT18",
            "gene_link": "https://www.uniprot.org/uniprot/Q6P9A2",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q6P9S7-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 10",
            "gene": "Galnt10",
            "gene_link": "https://www.uniprot.org/uniprot/Q6P9S7",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q6PB93-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 2",
            "gene": "Galnt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q6PB93",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q7TT15-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 17",
            "gene": "Galnt17",
            "gene_link": "https://www.uniprot.org/uniprot/Q7TT15",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q7Z4T8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Inactive polypeptide N-acetylgalactosaminyltransferase-like protein 5",
            "gene": "GALNTL5",
            "gene_link": "https://www.uniprot.org/uniprot/Q7Z4T8",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
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            ]
        },
        {
            "uniprot_canonical_ac": "Q7Z7M9-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 5",
            "gene": "GALNT5",
            "gene_link": "https://www.uniprot.org/uniprot/Q7Z7M9",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q80VA0-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "N-acetylgalactosaminyltransferase 7",
            "gene": "Galnt7",
            "gene_link": "https://www.uniprot.org/uniprot/Q80VA0",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q86SF2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
            "protein_name": "N-acetylgalactosaminyltransferase 7",
            "gene": "GALNT7",
            "gene_link": "https://www.uniprot.org/uniprot/Q86SF2",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00033MO",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00033MO"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q86SR1-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00033MO",
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