{
    "glytoucan_ac": "G03172PR",
    "number_monosaccharides": 5,
    "number_monosaccharides_suffix": "",
    "mass": 1168.41,
    "mass_pme": 1462.74,
    "tool_support": {
        "gnome_glygen_oglycans": "yes",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "yes",
        "pdb": "yes",
        "gnome_glygen_nglycans": "no",
        "gnome_glycotree_nglycans": "no"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b1-D-Neup5Ac-(2\u21926)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21923)[\u03b1-D-Neup5Ac-(2\u21926)]-\u03b1-D-GalpNAc-(1\u2192",
    "wurcs": "WURCS=2.0/4,5,4/[a2112h-1a_1-5_2*NCC/3=O][a2122h-1b_1-5_2*NCC/3=O][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O]/1-2-3-4-4/a3-b1_a6-e2_b4-c1_c6-d2",
    "glycoct": "RES 1b:a-dgal-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dgal-HEX-1:5 6b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 7s:n-acetyl 8b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 9s:n-acetyl LIN 1:1d(2+1)2n 2:1o(3+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(6+2)6d 6:6d(5+1)7n 7:1o(6+2)8d 8:8d(5+1)9n",
    "inchi": "InChI=1S/C44H72N4O32/c1-12(52)45-23-16(56)5-43(41(68)69,79-36(23)27(60)18(58)7-49)72-10-21-29(62)32(65)33(66)40(76-21)77-34-20(9-51)75-39(25(31(34)64)47-14(3)54)78-35-26(48-15(4)55)38(67)74-22(30(35)63)11-73-44(42(70)71)6-17(57)24(46-13(2)53)37(80-44)28(61)19(59)8-50/h16-40,49-51,56-67H,5-11H2,1-4H3,(H,45,52)(H,46,53)(H,47,54)(H,48,55)(H,68,69)(H,70,71)/t16-,17-,18+,19+,20+,21+,22+,23+,24+,25+,26+,27+,28+,29-,30-,31+,32-,33+,34+,35+,36+,37+,38-,39-,40-,43+,44+/m0/s1",
    "glycam": "DNeup5Aca2-6DGalpb1-4DGlcpNAcb1-3[DNeup5Aca2-6]DGalpNAca1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H](C[C@@](O[C@H]1[C@@H]([C@@H](CO)O)O)(C(=O)O)OC[C@@H]2[C@@H]([C@@H]([C@H]([C@H](O2)O)NC(=O)C)O[C@H]3[C@@H]([C@H]([C@@H]([C@H](O3)CO)O[C@H]4[C@@H]([C@H]([C@H]([C@H](O4)CO[C@@]5(C[C@@H]([C@H]([C@@H](O5)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)O)O)NC(=O)C)O)O",
    "inchi_key": {
        "key": "OGWSXYIUPTXQMU-GHZRNRJTSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/OGWSXYIUPTXQMU-GHZRNRJTSA-N"
    },
    "byonic": "HexNAc(2)Hex(1)NeuAc(2) % 1168.41296668",
    "gwb": "freeEnd--1a1D-GalNAc,p(--3b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuAc,p)--6a2D-NeuAc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "2822",
                    "url": "https://glyconnect.expasy.org/browser/structures/2822"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G03172PR",
                    "url": "https://glycosmos.org/glycans/show/G03172PR"
                },
                {
                    "database": "PubMed",
                    "id": "4066681",
                    "url": "https://glygen.org/publication/PubMed/4066681"
                }
            ],
            "common_name": "Human",
            "glygen_name": "Human",
            "reference_species": "Homo sapiens [9606]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "O-linked",
                "url": ""
            },
            "subtype": {
                "name": "Core 3",
                "url": ""
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "E5D8G1-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase",
            "gene": "Galntl6",
            "gene_link": "https://www.uniprot.org/uniprot/E5D8G1",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "G3X942-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase",
            "gene": "Galnt9",
            "gene_link": "https://www.uniprot.org/uniprot/G3X942",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O08832-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 4",
            "gene": "Galnt4",
            "gene_link": "https://www.uniprot.org/uniprot/O08832",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O08912-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 1",
            "gene": "Galnt1",
            "gene_link": "https://www.uniprot.org/uniprot/O08912",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P70419-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 3",
            "gene": "Galnt3",
            "gene_link": "https://www.uniprot.org/uniprot/P70419",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10471-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 2",
            "gene": "GALNT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q10471",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10472-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 1",
            "gene": "GALNT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q10472",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q14435-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 3",
            "gene": "GALNT3",
            "gene_link": "https://www.uniprot.org/uniprot/Q14435",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q49A17-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase-like 6",
            "gene": "GALNTL6",
            "gene_link": "https://www.uniprot.org/uniprot/Q49A17",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q6IS24-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 17",
            "gene": "GALNT17",
            "gene_link": "https://www.uniprot.org/uniprot/Q6IS24",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q6P9A2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 18",
            "gene": "GALNT18",
            "gene_link": "https://www.uniprot.org/uniprot/Q6P9A2",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q6P9S7-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 10",
            "gene": "Galnt10",
            "gene_link": "https://www.uniprot.org/uniprot/Q6P9S7",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q6PB93-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 2",
            "gene": "Galnt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q6PB93",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q7TT15-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 17",
            "gene": "Galnt17",
            "gene_link": "https://www.uniprot.org/uniprot/Q7TT15",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q7Z4T8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Inactive polypeptide N-acetylgalactosaminyltransferase-like protein 5",
            "gene": "GALNTL5",
            "gene_link": "https://www.uniprot.org/uniprot/Q7Z4T8",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q7Z7M9-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 5",
            "gene": "GALNT5",
            "gene_link": "https://www.uniprot.org/uniprot/Q7Z7M9",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q80VA0-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "N-acetylgalactosaminyltransferase 7",
            "gene": "Galnt7",
            "gene_link": "https://www.uniprot.org/uniprot/Q80VA0",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q86SF2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "N-acetylgalactosaminyltransferase 7",
            "gene": "GALNT7",
            "gene_link": "https://www.uniprot.org/uniprot/Q86SF2",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q86SR1-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 10",
            "gene": "GALNT10",
            "gene_link": "https://www.uniprot.org/uniprot/Q86SR1",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BGT9-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 12",
            "gene": "Galnt12",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BGT9",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BVG5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 14",
            "gene": "Galnt14",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BVG5",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8C102-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 5",
            "gene": "Galnt5",
            "gene_link": "https://www.uniprot.org/uniprot/Q8C102",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8C7U7-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 6",
            "gene": "Galnt6",
            "gene_link": "https://www.uniprot.org/uniprot/Q8C7U7",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8CF93-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 13",
            "gene": "Galnt13",
            "gene_link": "https://www.uniprot.org/uniprot/Q8CF93",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8IUC8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 13",
            "gene": "GALNT13",
            "gene_link": "https://www.uniprot.org/uniprot/Q8IUC8",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8IXK2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 12",
            "gene": "GALNT12",
            "gene_link": "https://www.uniprot.org/uniprot/Q8IXK2",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8K1B9-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 18",
            "gene": "Galnt18",
            "gene_link": "https://www.uniprot.org/uniprot/Q8K1B9",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G03172PR",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G03172PR"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8N3T1-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G03172PR",
            "protein_name": "Polypeptide N-acetylgalactosaminyltransferase 15",
            "gene": "GALNT15",
            "gene_link": "https://www.uniprot.org/uniprot/Q8N3T1",
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