{
    "glytoucan_ac": "G01856RL",
    "number_monosaccharides": 10,
    "number_monosaccharides_suffix": "",
    "mass": 1947.68,
    "mass_pme": 2438.23,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "no",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "\u03b1-D-Neup5Gc-(2\u21926)-\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)-\u03b1-D-Manp-(1\u21923)[\u03b2-D-Galp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21922)-\u03b1-D-Manp-(1\u21926)]-\u03b2-D-Manp-(1\u21924)-\u03b2-D-GlcpNAc-(1\u21924)-\u03b2-D-GlcpNAc-(1\u2192",
    "wurcs": "WURCS=2.0/5,10,9/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCCO/3=O]/1-1-2-3-1-4-5-3-1-4/a4-b1_b4-c1_c3-d1_c6-h1_d2-e1_e4-f1_f6-g2_h2-i1_i4-j1",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 11s:n-glycolyl 12b:a-dman-HEX-1:5 13b:b-dglc-HEX-1:5 14s:n-acetyl 15b:b-dgal-HEX-1:5 LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:9o(6+2)10d 10:10d(5+1)11n 11:5o(6+1)12d 12:12o(2+1)13d 13:13d(2+1)14n 14:13o(4+1)15d",
    "inchi": "InChI=1S/C73H121N5O55/c1-17(88)74-34-44(101)55(26(10-83)117-63(34)112)126-64-35(75-18(2)89)45(102)58(29(13-86)121-64)129-69-54(111)60(43(100)30(124-69)15-115-70-61(50(107)40(97)24(8-81)119-70)131-65-36(76-19(3)90)46(103)56(27(11-84)122-65)127-67-52(109)48(105)39(96)23(7-80)118-67)130-71-62(51(108)41(98)25(9-82)120-71)132-66-37(77-20(4)91)47(104)57(28(12-85)123-66)128-68-53(110)49(106)42(99)31(125-68)16-116-73(72(113)114)5-21(92)33(78-32(94)14-87)59(133-73)38(95)22(93)6-79/h21-31,33-71,79-87,92-93,95-112H,5-16H2,1-4H3,(H,74,88)(H,75,89)(H,76,90)(H,77,91)(H,78,94)(H,113,114)/t21-,22+,23+,24+,25+,26+,27+,28+,29+,30+,31+,33+,34+,35+,36+,37+,38+,39-,40+,41+,42-,43+,44+,45+,46+,47+,48-,49-,50-,51-,52+,53+,54-,55+,56+,57+,58+,59+,60-,61-,62-,63+,64-,65-,66-,67-,68-,69-,70-,71+,73+/m0/s1",
    "glycam": "",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H]([C@@H]([C@H](O[C@H]1O)CO)O[C@H]2[C@@H]([C@H]([C@@H]([C@H](O2)CO)O[C@H]3[C@H]([C@H]([C@@H]([C@H](O3)CO[C@@H]4[C@H]([C@H]([C@@H]([C@H](O4)CO)O)O)O[C@H]5[C@@H]([C@H]([C@@H]([C@H](O5)CO)O[C@H]6[C@@H]([C@H]([C@H]([C@H](O6)CO)O)O)O)O)NC(=O)C)O)O[C@@H]7[C@H]([C@H]([C@@H]([C@H](O7)CO)O)O)O[C@H]8[C@@H]([C@H]([C@@H]([C@H](O8)CO)O[C@H]9[C@@H]([C@H]([C@H]([C@H](O9)CO[C@@]1(C[C@@H]([C@H]([C@@H](O1)[C@@H]([C@@H](CO)O)O)NC(=O)CO)O)C(=O)O)O)O)O)O)NC(=O)C)O)O)NC(=O)C)O",
    "inchi_key": {
        "key": "QSTBYNZWODVAIS-LEONRSRXSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/QSTBYNZWODVAIS-LEONRSRXSA-N"
    },
    "byonic": "HexNAc(4)Hex(5)NeuGc(1) % 1947.68250388",
    "gwb": "freeEnd--1b1D-GlcNAc,p--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p--2b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuGc,p)--6a1D-Man,p--2b1D-GlcNAc,p--4b1D-Gal,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9913,
            "name": "Bos taurus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3007",
                    "url": "https://glyconnect.expasy.org/browser/structures/3007"
                },
                {
                    "database": "GlyCosmos",
                    "id": "G01856RL",
                    "url": "https://glycosmos.org/glycans/show/G01856RL"
                },
                {
                    "database": "PubMed",
                    "id": "3356193",
                    "url": "https://glygen.org/publication/PubMed/3356193"
                }
            ],
            "common_name": "Bovine",
            "glygen_name": "Bovine",
            "reference_species": "Bos taurus [9913]"
        },
        {
            "taxid": 9823,
            "name": "Sus scrofa",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "3007",
                    "url": "https://glyconnect.expasy.org/browser/structures/3007"
                },
                {
                    "database": "PubMed",
                    "id": "3356193",
                    "url": "https://glygen.org/publication/PubMed/3356193"
                }
            ],
            "common_name": "Pig",
            "glygen_name": "Pig",
            "reference_species": "Sus scrofa [9823]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Biantennary"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Complex",
                "url": ""
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10469-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q10469",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921V5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat2",
            "gene_link": "https://www.uniprot.org/uniprot/Q921V5",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15907-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 1",
            "gene": "ST6GAL1",
            "gene_link": "https://www.uniprot.org/uniprot/P15907",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q64685-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 1",
            "gene": "St6gal1",
            "gene_link": "https://www.uniprot.org/uniprot/Q64685",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G01856RL",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G01856RL"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q76K27-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G01856RL",
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 2",
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