{
    "uniprot_canonical_ac": "A0A0G2JYF9-1",
    "uniprot_ac": "A0A0G2JYF9",
    "uniprot_id": "A0A0G2JYF9_RAT",
    "mass": {
        "chemical_mass": 50068.0,
        "monoisotopic_mass": 50068.0
    },
    "refseq": {},
    "gene_names": [
        {
            "name": "Matn3",
            "resource": "UniProtKB",
            "type": "recommended",
            "url": "https://www.uniprot.org/uniprot/A0A0G2JYF9",
            "id": "A0A0G2JYF9"
        }
    ],
    "protein_names": [
        {
            "name": "Matrilin 3",
            "resource": "UniProtKB",
            "type": "synonym",
            "url": "",
            "id": "A0A0G2JYF9"
        }
    ],
    "function": [],
    "disease": [
        {
            "disease_id": "DOID:0070299",
            "recommended_name": {
                "id": "DOID:0070299",
                "resource": "Disease Ontology",
                "url": "http://disease-ontology.org/term/DOID:0070299",
                "name": "multiple epiphyseal dysplasia 5",
                "description": "A multiple epiphyseal dysplasia that has_material_basis_in heterozygous mutation in the MATN3 gene on chromosome 2p24."
            },
            "synonyms": [
                {
                    "id": "DOID:0070299",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:0070299",
                    "name": "BHMED",
                    "description": ""
                },
                {
                    "id": "DOID:0070299",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:0070299",
                    "name": "EDM5",
                    "description": ""
                },
                {
                    "id": "DOID:0070299",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:0070299",
                    "name": "bilateral hereditary microepiphyseal dysplasia",
                    "description": ""
                },
                {
                    "id": "DOID:0070299",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:0070299",
                    "name": "multiple epiphyseal dysplasia MATN3-related",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "multiple epiphyseal dysplasia type 5",
                    "description": "Multiple epiphyseal dysplasia type 5 is a multiple epiphyseal dysplasia characterized by an early-onset of pain and stiffness (involving knee and hip), progressive deformity of the extremities and precocious osteoarthritis associated with delayed and irregular ossification of epiphyses. Features specific to multiple epiphyseal dysplasia, type 5 include normal stature and lesser incidence of gait abnormalities. Radiographs reveal epiphyseal and metaphyseal irregularities. Multiple epiphyseal dysplasia type 5 follows an autosomal dominant mode of transmission."
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "BHMED",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "EDM5",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "MATN3 multiple epiphyseal dysplasia (disease)",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "MED5",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "Polyepiphyseal dysplasia type 5",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "bilateral hereditary micro-epiphyseal dysplasia",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "epiphyseal dysplasia, multiple, type 5",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "multiple epiphyseal dysplasia (disease) caused by mutation in MATN3",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "Microepiphyseal dysplasia, bilateral hereditary",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "epiphyseal dysplasia multiple 5",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "epiphyseal dysplasia, multiple, 5",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "multiple epiphyseal dysplasia 5",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "multiple epiphyseal dysplasia, MATN3 related",
                    "description": ""
                },
                {
                    "id": "MONDO:0011765",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0011765",
                    "name": "multiple epiphyseal dysplasia, MATN3-related",
                    "description": ""
                },
                {
                    "id": "MIM:607078",
                    "resource": "MIM",
                    "url": "https://www.omim.org/entry/607078",
                    "name": "Multiple epiphyseal dysplasia 5",
                    "description": "Multiple epiphyseal dysplasia 5"
                }
            ],
            "evidence": [
                {
                    "id": "0070299",
                    "database": "Alliance of Genome Resources",
                    "url": "https://www.alliancegenome.org/disease/DOID:0070299"
                }
            ]
        },
        {
            "disease_id": "DOID:2256",
            "recommended_name": {
                "id": "DOID:2256",
                "resource": "Disease Ontology",
                "url": "http://disease-ontology.org/term/DOID:2256",
                "name": "osteochondrodysplasia",
                "description": "A bone development disease that results_in defective development of cartilage or bone."
            },
            "synonyms": [
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "Cartilage Development disorder",
                    "description": ""
                },
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "Congenital anomaly of cartilage",
                    "description": ""
                },
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "Osteochondrodysplasia syndrome",
                    "description": ""
                },
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "chondrodystrophy",
                    "description": ""
                },
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "skeletal dysplasia",
                    "description": ""
                }
            ],
            "evidence": [
                {
                    "id": "2256",
                    "database": "Alliance of Genome Resources",
                    "url": "https://www.alliancegenome.org/disease/DOID:2256"
                }
            ]
        },
        {
            "disease_id": "DOID:8398",
            "recommended_name": {
                "id": "DOID:8398",
                "resource": "Disease Ontology",
                "url": "http://disease-ontology.org/term/DOID:8398",
                "name": "osteoarthritis",
                "description": "An arthritis that has_material_basis_in worn out cartilage located_in joint."
            },
            "synonyms": [
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "Osteoarthrosis and allied disorder",
                    "description": ""
                },
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "degenerative arthritis",
                    "description": ""
                },
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "degenerative joint disease",
                    "description": ""
                },
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "hypertrophic arthritis",
                    "description": ""
                },
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "osteoarthrosis",
                    "description": ""
                }
            ],
            "evidence": [
                {
                    "id": "8398",
                    "database": "Alliance of Genome Resources",
                    "url": "https://www.alliancegenome.org/disease/DOID:8398"
                }
            ]
        }
    ],
    "snv": [],
    "mutagenesis": [],
    "expression_tissue": [],
    "expression_disease": [],
    "species": [
        {
            "name": "Rattus norvegicus",
            "taxid": 10116,
            "evidence": [
                {
                    "database": "UniProtKB",
                    "id": "A0A0G2JYF9",
                    "url": "https://www.uniprot.org/uniprot/A0A0G2JYF9"
                }
            ],
            "common_name": "Rat"
        }
    ],
    "gene": [
        {
            "name": "Matn3",
            "url": "https://www.uniprot.org/uniprot/A0A0G2JYF9",
            "locus": {
                "chromosome": "6",
                "start_pos": 31768101,
                "end_pos": 31748474,
                "strand": "-",
                "evidence": [
                    {
                        "database": "Ensembl Gene",
                        "id": "ENSRNOG00000056141",
                        "url": "https://www.ensembl.org/id/ENSRNOG00000056141"
                    }
                ]
            }
        }
    ],
    "crossref": [
        {
            "id": "313954",
            "url": "https://www.ncbi.nlm.nih.gov/gene/313954",
            "database": "GeneID",
            "categories": [
                "Genome Annotation"
            ]
        },
        {
            "id": "IPR000742",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR000742",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR001881",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR001881",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR002035",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR002035",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR009030",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR009030",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR019466",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR019466",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR026823",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR026823",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR036465",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR036465",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "rno:313954",
            "url": "https://www.genome.jp/dbget-bin/get_linkdb?-t+pathway+rno:313954",
            "database": "KEGG Pathway",
            "categories": [
                "Enzyme and Pathway"
            ]
        },
        {
            "id": "CALGTHQ",
            "url": "https://omabrowser.org/oma/group/CALGTHQ/",
            "database": "OMA",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PTHR24020",
            "url": "http://www.pantherdb.org/panther/family.do?clsAccession=PTHR24020",
            "database": "PANTHER",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PTHR24020:SF12",
            "url": "http://www.pantherdb.org/panther/family.do?clsAccession=PTHR24020:SF12",
            "database": "PANTHER",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PF00092",
            "url": "http://pfam.xfam.org/family/PF00092",
            "database": "Pfam",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PF10393",
            "url": "http://pfam.xfam.org/family/PF10393",
            "database": "Pfam",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PF12662",
            "url": "http://pfam.xfam.org/family/PF12662",
            "database": "Pfam",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PF14670",
            "url": "http://pfam.xfam.org/family/PF14670",
            "database": "Pfam",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "R-RNO-3000178",
            "url": "https://www.reactome.org/PathwayBrowser/#R-RNO-3000178",
            "database": "Reactome",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "R-RNO-381426",
            "url": "https://www.reactome.org/PathwayBrowser/#R-RNO-381426",
            "database": "Reactome",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "R-RNO-8957275",
            "url": "https://www.reactome.org/PathwayBrowser/#R-RNO-8957275",
            "database": "Reactome",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "NP_001101483",
            "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001101483",
            "database": "RefSeq",
            "categories": [
                "Protein"
            ]
        },
        {
            "id": "1305085",
            "url": "http://rgd.mcw.edu/tools/genes/genes_view.cgi?id=1305085",
            "database": "RGD",
            "categories": [
                "Organism-specific"
            ]
        },
        {
            "id": "A0A0G2JYF9",
            "url": "https://www.uniprot.org/uniprot/A0A0G2JYF9",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        },
        {
            "id": "A0A0G2JYF9-1",
            "url": "https://www.uniprot.org/uniprot/A0A0G2JYF9-1",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        }
    ],
    "sequence": {
        "sequence": "MLFSALLRNLPGLLLLLWPLLLLPPPAAPGRLTRASVRRLGTRVPGGSPGHFSARATSTRAPYSGVRGSGVCKSRPLDLVFIIDSSRSVRPLEFTKVKTFVSRIIDTLDIGATDTRVAVVNYASTVKIEFQLNTYSNKQALKQAVARITPLSTGTMSGLAIQTAMEEAFTVEAGARGPTSNIPKVAIIVTDGRPQDQVNEVAARARASGIELYAVGVDRADMESLKMMASKPLEDHVFYVETYGVIEKLSARFQETFCALDQCMLGTHQCQHVCVSDGDGRHHCECSQGYTLNADGKTCSAIDKCALNTHGCEQICVNDRNGSYHCECYEGYTLNADRKTCAAQDKCALGTHGCQHICVNDGAGSHHCECFEGYTLNADKKTCSVRDKCALGTHGCQHICVSDGAVSYHCDCFPGYTLNDDKKTCSDIEEVRSLISIEDACSCGATLAFQEKVSSHLQKLNTT",
        "length": 463,
        "header": "tr|A0A0G2JYF9-1|A0A0G2JYF9_RAT Isoform 1 of Matrilin 3  OS=Rattus norvegicus OX=10116 GN=Matn3 PE=4 SV=2 CANONICAL=A0A0G2JYF9-1"
    },
    "isoforms": [
        {
            "isoform_ac": "A0A0G2JYF9-1",
            "url": "",
            "sequence": {
                "sequence": "MLFSALLRNLPGLLLLLWPLLLLPPPAAPGRLTRASVRRLGTRVPGGSPGHFSARATSTRAPYSGVRGSGVCKSRPLDLVFIIDSSRSVRPLEFTKVKTFVSRIIDTLDIGATDTRVAVVNYASTVKIEFQLNTYSNKQALKQAVARITPLSTGTMSGLAIQTAMEEAFTVEAGARGPTSNIPKVAIIVTDGRPQDQVNEVAARARASGIELYAVGVDRADMESLKMMASKPLEDHVFYVETYGVIEKLSARFQETFCALDQCMLGTHQCQHVCVSDGDGRHHCECSQGYTLNADGKTCSAIDKCALNTHGCEQICVNDRNGSYHCECYEGYTLNADRKTCAAQDKCALGTHGCQHICVNDGAGSHHCECFEGYTLNADKKTCSVRDKCALGTHGCQHICVSDGAVSYHCDCFPGYTLNDDKKTCSDIEEVRSLISIEDACSCGATLAFQEKVSSHLQKLNTT",
                "length": 463,
                "header": "tr|A0A0G2JYF9-1|A0A0G2JYF9_RAT Isoform 1 of Matrilin 3  OS=Rattus norvegicus OX=10116 GN=Matn3 PE=4 SV=2 CANONICAL=A0A0G2JYF9-1"
            },
            "locus": {
                "chromosome": "6",
                "start_pos": 31767601,
                "end_pos": 31748587,
                "strand": "-",
                "evidence": [
                    {
                        "database": "Ensembl Transcript",
                        "id": "ENSRNOT00000084212.2",
                        "url": "https://www.ensembl.org/id/ENSRNOT00000084212.2"
                    },
                    {
                        "database": "Ensembl Peptide",
                        "id": "ENSRNOP00000070640.2",
                        "url": "https://www.ensembl.org/id/ENSRNOP00000070640.2"
                    }
                ]
            }
        }
    ],
    "glycosylation": [],
    "phosphorylation": [],
    "glycation": [],
    "keywords": [],
    "publication": [],
    "pathway": [
        {
            "id": "rno:313954",
            "name": "",
            "url": "https://www.genome.jp/dbget-bin/get_linkdb?-t+pathway+rno:313954",
            "resource": "KEGG Pathway"
        },
        {
            "id": "R-RNO-3000178",
            "name": "ECM proteoglycans",
            "url": "https://www.reactome.org/PathwayBrowser/#R-RNO-3000178",
            "resource": "Reactome",
            "description": "ECM proteoglycans"
        },
        {
            "id": "R-RNO-381426",
            "name": "Regulation of Insulin-like Growth Factor (IGF) transport and uptake by Insulin-like Growth Factor Binding Proteins (IGFBPs)",
            "url": "https://www.reactome.org/PathwayBrowser/#R-RNO-381426",
            "resource": "Reactome",
            "description": "Regulation of Insulin-like Growth Factor (IGF) transport and uptake by Insulin-like Growth Factor Binding Proteins (IGFBPs)"
        },
        {
            "id": "R-RNO-8957275",
            "name": "Post-translational protein phosphorylation",
            "url": "https://www.reactome.org/PathwayBrowser/#R-RNO-8957275",
            "resource": "Reactome",
            "description": "Post-translational protein phosphorylation"
        }
    ],
    "orthologs": [
        {
            "uniprot_canonical_ac": "O15232-1",
            "protein_name": "Matrilin-3",
            "gene_name": "MATN3",
            "tax_id": 9606,
            "organism": "Homo sapiens",
            "sequence": {
                "sequence": "MPRPAPARRLPGLLLLLWPLLLLPSAAPDPVARPGFRRLETRGPGGSPGRRPSPAAPDGAPASGTSEPGRARGAGVCKSRPLDLVFIIDSSRSVRPLEFTKVKTFVSRIIDTLDIGPADTRVAVVNYASTVKIEFQLQAYTDKQSLKQAVGRITPLSTGTMSGLAIQTAMDEAFTVEAGAREPSSNIPKVAIIVTDGRPQDQVNEVAARAQASGIELYAVGVDRADMASLKMMASEPLEEHVFYVETYGVIEKLSSRFQETFCALDPCVLGTHQCQHVCISDGEGKHHCECSQGYTLNADKKTCSALDRCALNTHGCEHICVNDRSGSYHCECYEGYTLNEDRKTCSAQDKCALGTHGCQHICVNDRTGSHHCECYEGYTLNADKKTCSVRDKCALGSHGCQHICVSDGAASYHCDCYPGYTLNEDKKTCSATEEARRLVSTEDACGCEATLAFQDKVSSYLQRLNTKLDDILEKLKINEYGQIHR",
                "length": 486
            },
            "evidence": [
                {
                    "id": "45226888",
                    "url": "http://www.informatics.jax.org/homology/45226888",
                    "database": "MGI"
                }
            ],
            "common_name": "Human"
        },
        {
            "uniprot_canonical_ac": "O35701-1",
            "protein_name": "Matrilin-3",
            "gene_name": "Matn3",
            "tax_id": 10090,
            "organism": "Mus musculus",
            "sequence": {
                "sequence": "MLLSAPLRHLPGLLLLLWPLLLLPSLAAPGRLARASVRRLGTRVPGGSPGHLSALATSTRAPYSGGRGAGVCKSRPLDLVFIIDSSRSVRPLEFTKVKTFVSRIIDTLDIGATDTRVAVVNYASTVKIEFQLNTYSDKQALKQAVARITPLSTGTMSGLAIQTAMEEAFTVEAGARGPMSNIPKVAIIVTDGRPQDQVNEVAARARASGIELYAVGVDRADMESLKMMASKPLEEHVFYVETYGVIEKLSARFQETFCALDQCMLGTHQCQHVCVSDGDGKHHCECSQGYTLNADGKTCSAIDKCALSTHGCEQICVNDRNGSYHCECYGGYALNADRRTCAALDKCASGTHGCQHICVNDGAGSHHCECFEGYTLNADKKTCSVRNKCALGTHGCQHICVSDGAVAYHCDCFPGYTLNDDKKTCSDIEEARSLISIEDACGCGATLAFQEKVSSHLQKLNTKLDNILKKLKVTEYGQVHR",
                "length": 481
            },
            "evidence": [
                {
                    "id": "45226888",
                    "url": "http://www.informatics.jax.org/homology/45226888",
                    "database": "MGI"
                }
            ],
            "common_name": "Mouse"
        }
    ],
    "go_annotation": {
        "categories": [
            {
                "name": "Biological Process",
                "total": 2,
                "go_terms": [
                    {
                        "name": "extracellular matrix organization",
                        "id": "GO:0030198",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0030198",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "A0A0G2JYF9",
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                        "pmid": ""
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                "name": "Cellular Component",
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            {
                "name": "Molecular Function",
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                        "name": "calcium ion binding",
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                        "pmid": ""
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    "pro_annotation": [],
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            "start_pos": 321,
            "end_pos": 323,
            "annotation": "n_glycosylation_sequon"
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            "start_pos": 461,
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    "ptm_annotation": [],
    "interactions": [],
    "reactions": [],
    "reaction_participants": [],
    "reaction_enzymes": [],
    "biomarkers": [],
    "synthesized_glycans": [],
    "cluster_types": [
        {
            "name": "homologset.mgi",
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        {
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