{
    "uniprot_canonical_ac": "Q5I0M4-1",
    "uniprot_ac": "Q5I0M4",
    "uniprot_id": "Q5I0M4_RAT",
    "mass": {
        "chemical_mass": 37013.0,
        "monoisotopic_mass": 37013.0
    },
    "refseq": {
        "ac": "NP_001014262",
        "name": "aldo-keto reductase family 1, member C12",
        "summary": "",
        "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262"
    },
    "gene_names": [
        {
            "name": "Akr1c12",
            "resource": "UniProtKB",
            "type": "recommended",
            "url": "https://www.uniprot.org/uniprot/Q5I0M4",
            "id": "Q5I0M4"
        },
        {
            "name": "Akr1c12",
            "resource": "RefSeq",
            "type": "synonym",
            "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262",
            "id": "NP_001014262"
        },
        {
            "name": "Akr1c13; BER1; LOCT364773; TBER1",
            "resource": "RefSeq",
            "type": "synonym",
            "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262",
            "id": "NP_001014262"
        }
    ],
    "protein_names": [
        {
            "name": "Aldo-keto reductase family 1, member C12",
            "resource": "UniProtKB",
            "type": "synonym",
            "url": "",
            "id": "Q5I0M4"
        },
        {
            "name": "aldo-keto reductase family 1, member C12",
            "resource": "RefSeq",
            "type": "synonym",
            "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262",
            "id": "NP_001014262"
        },
        {
            "name": "liver regeneration-related protein LRRG07; 17beta-hydroxysteroid dehydrogenase; aldo-keto reductase family 1, member C13",
            "resource": "RefSeq",
            "type": "synonym",
            "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262",
            "id": "NP_001014262"
        }
    ],
    "function": [
        {
            "annotation": "Analysis of rat and mouse NAD+-dependent 3alpha/17beta/20alpha-hydroxysteroid dehydrogenases.",
            "evidence": [
                {
                    "database": "RefSeq",
                    "id": "NP_001014262",
                    "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262"
                },
                {
                    "database": "PubMed",
                    "id": "17888864",
                    "url": "https://glygen.org/publication/PubMed/17888864"
                }
            ]
        },
        {
            "annotation": "TBER1 represents a novel type of 17beta-hydroxysteroid dehydrogenase with unique catalytic properties and tissue distribution that is distinct from the type 5 isozyme.",
            "evidence": [
                {
                    "database": "RefSeq",
                    "id": "NP_001014262",
                    "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262"
                },
                {
                    "database": "PubMed",
                    "id": "16788056",
                    "url": "https://glygen.org/publication/PubMed/16788056"
                }
            ]
        }
    ],
    "disease": [
        {
            "disease_id": "DOID:0111773",
            "recommended_name": {
                "id": "DOID:0111773",
                "resource": "Disease Ontology",
                "url": "http://disease-ontology.org/term/DOID:0111773",
                "name": "46,XY sex reversal 8",
                "description": "A 46 XY sex reversal characterized by an XY karyotype, phenotypically female genitalia and failure to develop secondary sexual characteristics at puberty including menstruation that has_material_basis_in homozygous or compound heterozygous mutation in the AKR1C2 gene on chromosome 10p15.1."
            },
            "synonyms": [
                {
                    "id": "DOID:0111773",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:0111773",
                    "name": "SRXY8",
                    "description": ""
                },
                {
                    "id": "DOID:0111773",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:0111773",
                    "name": "TDD",
                    "description": ""
                },
                {
                    "id": "DOID:0111773",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:0111773",
                    "name": "male pseudohermaphroditism due to deficiency of testicular 17,20-desmolase",
                    "description": ""
                },
                {
                    "id": "MONDO:0013664",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013664",
                    "name": "46,XY disorder of sex development due to testicular 17,20-desmolase deficiency",
                    "description": ""
                },
                {
                    "id": "MONDO:0013664",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013664",
                    "name": "46,XY sex reversal type 8",
                    "description": ""
                },
                {
                    "id": "MONDO:0013664",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013664",
                    "name": "46XY sex reversal 8",
                    "description": ""
                },
                {
                    "id": "MONDO:0013664",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013664",
                    "name": "46XY sex reversal 8, modifier of",
                    "description": ""
                },
                {
                    "id": "MONDO:0013664",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013664",
                    "name": "46,XY sex reversal 8",
                    "description": ""
                },
                {
                    "id": "MONDO:0013664",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013664",
                    "name": "SRXY8",
                    "description": ""
                },
                {
                    "id": "MONDO:0013664",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013664",
                    "name": "male pseudohermaphroditism due to deficiency of testicular 17,20-desmolase",
                    "description": ""
                },
                {
                    "id": "MIM:614279",
                    "resource": "MIM",
                    "url": "https://www.omim.org/entry/614279",
                    "name": "46,XY sex reversal 8",
                    "description": "46,XY sex reversal 8"
                }
            ],
            "evidence": [
                {
                    "id": "0111773",
                    "database": "Alliance of Genome Resources",
                    "url": "https://www.alliancegenome.org/disease/DOID:0111773"
                }
            ]
        }
    ],
    "snv": [],
    "mutagenesis": [],
    "expression_tissue": [],
    "expression_disease": [],
    "species": [
        {
            "name": "Rattus norvegicus",
            "taxid": 10116,
            "evidence": [
                {
                    "database": "UniProtKB",
                    "id": "Q5I0M4",
                    "url": "https://www.uniprot.org/uniprot/Q5I0M4"
                }
            ],
            "common_name": "Rat"
        }
    ],
    "gene": [
        {
            "name": "Akr1c12",
            "url": "https://www.uniprot.org/uniprot/Q5I0M4",
            "locus": {
                "chromosome": "17",
                "start_pos": 66045378,
                "end_pos": 66108329,
                "strand": "+",
                "evidence": [
                    {
                        "database": "Ensembl Gene",
                        "id": "ENSRNOG00000033010",
                        "url": "https://www.ensembl.org/id/ENSRNOG00000033010"
                    }
                ]
            }
        }
    ],
    "crossref": [
        {
            "id": "ENSRNOG00000033010",
            "url": "https://bgee.org/?page=gene&gene_id=ENSRNOG00000033010",
            "database": "Bgee",
            "categories": [
                "Gene Expression"
            ]
        },
        {
            "id": "cd19108",
            "url": "https://www.ncbi.nlm.nih.gov/Structure/cdd/cddsrv.cgi?uid=cd19108",
            "database": "CDD",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "364773",
            "url": "https://www.ncbi.nlm.nih.gov/gene/364773",
            "database": "GeneID",
            "categories": [
                "Genome Annotation"
            ]
        },
        {
            "id": "IPR018170",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR018170",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR020471",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR020471",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR023210",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR023210",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR036812",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR036812",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR044482",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR044482",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "rno:364773",
            "url": "https://www.genome.jp/dbget-bin/get_linkdb?-t+pathway+rno:364773",
            "database": "KEGG Pathway",
            "categories": [
                "Enzyme and Pathway"
            ]
        },
        {
            "id": "VHYPLAT",
            "url": "https://omabrowser.org/oma/group/VHYPLAT/",
            "database": "OMA",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "890110at2759",
            "url": "http://www.orthodb.org/?query=890110at2759",
            "database": "OrthoDB",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PTHR11732",
            "url": "http://www.pantherdb.org/panther/family.do?clsAccession=PTHR11732",
            "database": "PANTHER",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PTHR11732:SF286",
            "url": "http://www.pantherdb.org/panther/family.do?clsAccession=PTHR11732:SF286",
            "database": "PANTHER",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PF00248",
            "url": "http://pfam.xfam.org/family/PF00248",
            "database": "Pfam",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "NP_001014262",
            "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262",
            "database": "RefSeq",
            "categories": [
                "Protein"
            ]
        },
        {
            "id": "1359406",
            "url": "http://rgd.mcw.edu/tools/genes/genes_view.cgi?id=1359406",
            "database": "RGD",
            "categories": [
                "Organism-specific"
            ]
        },
        {
            "id": "Q5I0M4",
            "url": "https://www.uniprot.org/uniprot/Q5I0M4",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        },
        {
            "id": "Q5I0M4-1",
            "url": "https://www.uniprot.org/uniprot/Q5I0M4-1",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        }
    ],
    "sequence": {
        "sequence": "MSSKLHCVKLNDGHFIPALGFGTYKPKEVPKSKSLEAAHLAIDVGYRHIDTASAYQVEEEIGQAIQSKIKAGVVKRKDMFITTKLWCSCFRTEMVRPALEKSLKNLQLDYVDLFLIHYPVPIKSSVDESPLDEKGKFLLDTVDFCDTWEMLEKCKDAGLVKSIGVSNFNHKQLERLLNKPGLKYKPVCNQVECHLYLNQSKLLDYCKSKDIVLVAYGALGTQRYKEWVDQNSPVLLDDPILCDVAKKNKRSPALIALRYLFQRGVVPLAQSFKENEMRENLQVFEFQLSPEDMKTLDGLNKNFRYLSAEFLADHPEYPFSEEY",
        "length": 323,
        "header": "tr|Q5I0M4-1|Q5I0M4_RAT Isoform 1 of Aldo-keto reductase family 1, member C12  OS=Rattus norvegicus OX=10116 GN=Akr1c12 PE=3 SV=1 CANONICAL=Q5I0M4-1"
    },
    "isoforms": [
        {
            "isoform_ac": "Q5I0M4-1",
            "url": "",
            "sequence": {
                "sequence": "MSSKLHCVKLNDGHFIPALGFGTYKPKEVPKSKSLEAAHLAIDVGYRHIDTASAYQVEEEIGQAIQSKIKAGVVKRKDMFITTKLWCSCFRTEMVRPALEKSLKNLQLDYVDLFLIHYPVPIKSSVDESPLDEKGKFLLDTVDFCDTWEMLEKCKDAGLVKSIGVSNFNHKQLERLLNKPGLKYKPVCNQVECHLYLNQSKLLDYCKSKDIVLVAYGALGTQRYKEWVDQNSPVLLDDPILCDVAKKNKRSPALIALRYLFQRGVVPLAQSFKENEMRENLQVFEFQLSPEDMKTLDGLNKNFRYLSAEFLADHPEYPFSEEY",
                "length": 323,
                "header": "tr|Q5I0M4-1|Q5I0M4_RAT Isoform 1 of Aldo-keto reductase family 1, member C12  OS=Rattus norvegicus OX=10116 GN=Akr1c12 PE=3 SV=1 CANONICAL=Q5I0M4-1"
            },
            "locus": {
                "chromosome": "17",
                "start_pos": 66061467,
                "end_pos": 66045595,
                "strand": "+",
                "evidence": [
                    {
                        "database": "Ensembl Transcript",
                        "id": "ENSRNOT00000117319.1",
                        "url": "https://www.ensembl.org/id/ENSRNOT00000117319.1"
                    },
                    {
                        "database": "Ensembl Peptide",
                        "id": "ENSRNOP00000097567.1",
                        "url": "https://www.ensembl.org/id/ENSRNOP00000097567.1"
                    }
                ]
            }
        }
    ],
    "glycosylation": [],
    "phosphorylation": [],
    "glycation": [],
    "keywords": [],
    "publication": [
        {
            "title": "Molecular cloning of a novel type of rat cytoplasmic 17beta-hydroxysteroid dehydrogenase distinct from the type 5 isozyme.",
            "journal": "Journal of biochemistry",
            "date": "2006",
            "authors": "Ishikura S, Matsumoto K, Sanai M, Horie K, Matsunaga T, Tajima K, El-Kabbani O, Hara A",
            "evidence": [
                {
                    "database": "RefSeq",
                    "id": "NP_001014262",
                    "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262"
                }
            ],
            "reference": [
                {
                    "type": "PubMed",
                    "id": "16788056",
                    "url": "https://glygen.org/publication/PubMed/16788056"
                }
            ]
        },
        {
            "title": "Characterization of rat and mouse NAD+-dependent 3alpha/17beta/20alpha-hydroxysteroid dehydrogenases and identification of substrate specificity determinants by site-directed mutagenesis.",
            "journal": "Archives of biochemistry and biophysics",
            "date": "2007",
            "authors": "Endo S, Sanai M, Horie K, Matsunaga T, Ishikura S, Tajima K, El-Kabbani O, Hara A",
            "evidence": [
                {
                    "database": "RefSeq",
                    "id": "NP_001014262",
                    "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001014262"
                }
            ],
            "reference": [
                {
                    "type": "PubMed",
                    "id": "17888864",
                    "url": "https://glygen.org/publication/PubMed/17888864"
                }
            ]
        }
    ],
    "pathway": [
        {
            "id": "rno:364773",
            "name": "",
            "url": "https://www.genome.jp/dbget-bin/get_linkdb?-t+pathway+rno:364773",
            "resource": "KEGG Pathway"
        }
    ],
    "orthologs": [
        {
            "uniprot_canonical_ac": "A0A8I6AD50-1",
            "protein_name": "Aldo-keto reductase family 1, member C13",
            "gene_name": "Akr1c13",
            "tax_id": 10116,
            "organism": "Rattus norvegicus",
            "sequence": {
                "sequence": "MAASCHYWAMALFKIQSSEYVNQISFGQAEYVAYLQRAEGRMSPLSEPEQAAGFLQVMRHCSCCLCSPGVSASYKLVPKSKSLEAAHLAIDAGYRHIDTAYAYQIEEEIGQAIQSKIKAGVVKREDMFITTKLWCTCFRPELVKPALEKSLKNLQLDYADLYIMHYPVPMKSGDNDFPVDEKGKSLLDTVDFCDTWEMLEKCKDAGLVKSIGVSNFNHKQLERLLNKPGLKYKPVCNQVECHLYLNQSKLLDYCKSKDIVLVAYGALGTQRYKEWVDQNSPVLLNDPVLCDVAKKNKRSPALIALRYLVQRGVVPLAQSFKENEMRENLQVFDFQLSPEDMKTLDGLNKNFRYLSAEFLAGHPEYPFSEEY",
                "length": 371
            },
            "evidence": [
                {
                    "id": "45240331",
                    "url": "http://www.informatics.jax.org/homology/45240331",
                    "database": "MGI"
                },
                {
                    "id": "45240330",
                    "url": "http://www.informatics.jax.org/homology/45240330",
                    "database": "MGI"
                }
            ],
            "common_name": "Rat"
        },
        {
            "uniprot_canonical_ac": "A2VD16-1",
            "protein_name": "Aldo-keto reductase family 1, member C12-like 1",
            "gene_name": "Akr1c12l1",
            "tax_id": 10116,
            "organism": "Rattus norvegicus",
            "sequence": {
                "sequence": "MSSKHHCVKLNDGHFIPALGFGTSIPNEVPKSKSLEAVHLAIDAGYHHIDTASAYQIEEEIGQAIQSKIKAGVVKREDMFITTKLWCTCFRPELVKPALEKSLKNLQLDYADLYIMHYPVPMKSGDKYLPVDDKGKWLLDTVDFCDTWEMLEKCKDAGLVKSIGVSNFNHKQLERLLNKPGLKYKPVCNQVECHLYMNQSKLLDYCKSKDIVLVAYGALGTQRYKEWVDQNSPVLLDDPVLCDVAKKNKRSPALIALRYLVQREVVPLAQSFKENEMRENLQVFEFQLSPEDMKTLDGLNKNFRYLSAEFLAGHPEYPFSEEY",
                "length": 323
            },
            "evidence": [
                {
                    "id": "45240331",
                    "url": "http://www.informatics.jax.org/homology/45240331",
                    "database": "MGI"
                },
                {
                    "id": "45240330",
                    "url": "http://www.informatics.jax.org/homology/45240330",
                    "database": "MGI"
                }
            ],
            "common_name": "Rat"
        },
        {
            "uniprot_canonical_ac": "P17516-1",
            "protein_name": "Aldo-keto reductase family 1 member C4",
            "gene_name": "AKR1C4",
            "tax_id": 9606,
            "organism": "Homo sapiens",
            "sequence": {
                "sequence": "MDPKYQRVELNDGHFMPVLGFGTYAPPEVPRNRAVEVTKLAIEAGFRHIDSAYLYNNEEQVGLAIRSKIADGSVKREDIFYTSKLWCTFFQPQMVQPALESSLKKLQLDYVDLYLLHFPMALKPGETPLPKDENGKVIFDTVDLSATWEVMEKCKDAGLAKSIGVSNFNCRQLEMILNKPGLKYKPVCNQVECHPYLNQSKLLDFCKSKDIVLVAHSALGTQRHKLWVDPNSPVLLEDPVLCALAKKHKQTPALIALRYQLQRGVVVLAKSYNEQRIRENIQVFEFQLTSEDMKVLDGLNRNYRYVVMDFLMDHPDYPFSDEY",
                "length": 323
            },
            "evidence": [
                {
                    "id": "45240331",
                    "url": "http://www.informatics.jax.org/homology/45240331",
                    "database": "MGI"
                },
                {
                    "id": "45240330",
                    "url": "http://www.informatics.jax.org/homology/45240330",
                    "database": "MGI"
                }
            ],
            "common_name": "Human"
        },
        {
            "uniprot_canonical_ac": "Q8VC28-1",
            "protein_name": "Aldo-keto reductase family 1 member C13",
            "gene_name": "Akr1c13",
            "tax_id": 10090,
            "organism": "Mus musculus",
            "sequence": {
                "sequence": "MSSKQHCVKLNDGHLIPALGFGTYKPKEVPKSKSLEAACLALDVGYRHVDTAYAYQVEEEIGQAIQSKIKAGVVKREDLFITTKLWCTCFRPELVKPALEKSLKKLQLDYVDLYIMHYPVPMKSGDNDFPVNEQGKSLLDTVDFCDTWERLEECKDAGLVKSIGVSNFNHRQLERILNKPGLKYKPVCNQVECHLYLNQRKLLDYCESKDIVLVAYGALGTQRYKEWVDQNSPVLLNDPVLCDVAKKNKRSPALIALRYLIQRGIVPLAQSFKENEMRENLQVFGFQLSPEDMKTLDGLNKNFRYLPAEFLVDHPEYPFVEEY",
                "length": 323
            },
            "evidence": [
                {
                    "id": "45240331",
                    "url": "http://www.informatics.jax.org/homology/45240331",
                    "database": "MGI"
                }
            ],
            "common_name": "Mouse"
        },
        {
            "uniprot_canonical_ac": "Q9JLI0-1",
            "protein_name": "Aldo-keto reductase a",
            "gene_name": "Akr1c12",
            "tax_id": 10090,
            "organism": "Mus musculus",
            "sequence": {
                "sequence": "MSSKQHYVKLNDGHLIPALGFGTYKPKEVPKSKSLEAACLALDVGYRHVDTAYAYQVEEEIGQAIQSKIKAGVVKREDLFITTKLWCGCFRPELVKPALEKSLKSLQLDYVDLYLIHYPVPMKPGDNESPLDENGKFLLDTVDFCDTWERLEECKDAGLVKSIGVSNFNHRQLERILNKPGLKYKPVCNQVECHLYLNQSKLLDYCKSKDIVLVAYGALGTQRYKEWVDQNSPVLLNDPVLCDVAKRNKRSPALIALRYLFQRGIVPLAQSFKENEMRENLQVFEFQLSPEDMKTLDGLNKNFRYLPAEFLADHPEYPFSEEY",
                "length": 323
            },
            "evidence": [
                {
                    "id": "45240330",
                    "url": "http://www.informatics.jax.org/homology/45240330",
                    "database": "MGI"
                }
            ],
            "common_name": "Mouse"
        }
    ],
    "go_annotation": {
        "categories": [
            {
                "name": "Molecular Function",
                "total": 3,
                "go_terms": [
                    {
                        "name": "steroid dehydrogenase activity",
                        "id": "GO:0016229",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0016229",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "Q5I0M4",
                                "url": "https://www.uniprot.org/uniprot/Q5I0M4"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "alditol:NADP+ 1-oxidoreductase activity",
                        "id": "GO:0004032",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0004032",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "Q5I0M4",
                                "url": "https://www.uniprot.org/uniprot/Q5I0M4"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "ketosteroid monooxygenase activity",
                        "id": "GO:0047086",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0047086",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "Q5I0M4",
                                "url": "https://www.uniprot.org/uniprot/Q5I0M4"
                            }
                        ],
                        "pmid": ""
                    }
                ]
            },
            {
                "name": "Cellular Component",
                "total": 1,
                "go_terms": [
                    {
                        "name": "cytosol",
                        "id": "GO:0005829",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0005829",
                        "evidence": [
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                                "database": "UniProtKB",
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