{
    "uniprot_canonical_ac": "P20850-1",
    "uniprot_ac": "P20850",
    "uniprot_id": "CO9A1_RAT",
    "mass": {
        "chemical_mass": 31256.0,
        "monoisotopic_mass": 31256.0
    },
    "refseq": {
        "ac": "NP_001094312",
        "name": "collagen alpha-1(IX) chain",
        "summary": "",
        "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001094312"
    },
    "gene_names": [
        {
            "name": "Col9a1",
            "resource": "UniProtKB",
            "type": "recommended",
            "url": "https://www.uniprot.org/uniprot/P20850",
            "id": "P20850"
        },
        {
            "name": "Col9a1",
            "resource": "RefSeq",
            "type": "synonym",
            "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001094312",
            "id": "NP_001094312"
        }
    ],
    "protein_names": [
        {
            "name": "Collagen alpha-1(IX) chain",
            "resource": "UniProtKB",
            "type": "recommended",
            "url": "",
            "id": "P20850"
        },
        {
            "name": "collagen alpha-1(IX) chain",
            "resource": "RefSeq",
            "type": "synonym",
            "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001094312",
            "id": "NP_001094312"
        },
        {
            "name": "procollagen, type IX, alpha 1; collagen, type IX, alpha 1",
            "resource": "RefSeq",
            "type": "synonym",
            "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001094312",
            "id": "NP_001094312"
        }
    ],
    "function": [
        {
            "annotation": "Collagen type IX alpha1 was found within the tectorial membrane as well as fibrocytes in the spiral ligament.",
            "evidence": [
                {
                    "database": "RefSeq",
                    "id": "NP_001094312",
                    "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001094312"
                },
                {
                    "database": "PubMed",
                    "id": "18448257",
                    "url": "https://glygen.org/publication/PubMed/18448257"
                }
            ]
        },
        {
            "annotation": "Structural component of hyaline cartilage and vitreous of the eye.",
            "evidence": [
                {
                    "database": "UniProtKB",
                    "id": "P20850",
                    "url": "https://www.uniprot.org/uniprot/P20850#function"
                }
            ]
        }
    ],
    "disease": [
        {
            "disease_id": "DOID:0070301",
            "recommended_name": {
                "id": "DOID:0070301",
                "resource": "Disease Ontology",
                "url": "http://disease-ontology.org/term/DOID:0070301",
                "name": "multiple epiphyseal dysplasia 6",
                "description": "A multiple epiphyseal dysplasia due to collagen 9 anomaly that has_material_basis_in heterozygous mutation in the COL9A1 gene on chromosome 6p13."
            },
            "synonyms": [
                {
                    "id": "DOID:0070301",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:0070301",
                    "name": "EDM6",
                    "description": ""
                },
                {
                    "id": "MONDO:0013591",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013591",
                    "name": "epiphyseal dysplasia, multiple, 6",
                    "description": "Any multiple epiphyseal dysplasia in which the cause of the disease is a mutation in the COL9A1 gene."
                },
                {
                    "id": "MONDO:0013591",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013591",
                    "name": "COL9A1 multiple epiphyseal dysplasia (disease)",
                    "description": ""
                },
                {
                    "id": "MONDO:0013591",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013591",
                    "name": "epiphyseal dysplasia, multiple, 6",
                    "description": ""
                },
                {
                    "id": "MONDO:0013591",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013591",
                    "name": "epiphyseal dysplasia, multiple, type 6",
                    "description": ""
                },
                {
                    "id": "MONDO:0013591",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013591",
                    "name": "multiple epiphyseal dysplasia (disease) caused by mutation in COL9A1",
                    "description": ""
                },
                {
                    "id": "MONDO:0013591",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013591",
                    "name": "EDM6",
                    "description": ""
                },
                {
                    "id": "MONDO:0013591",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0013591",
                    "name": "multiple epiphyseal dysplasia 6",
                    "description": ""
                },
                {
                    "id": "MIM:614135",
                    "resource": "MIM",
                    "url": "https://www.omim.org/entry/614135",
                    "name": "Multiple epiphyseal dysplasia 6",
                    "description": "Multiple epiphyseal dysplasia 6"
                }
            ],
            "evidence": [
                {
                    "id": "0070301",
                    "database": "Alliance of Genome Resources",
                    "url": "https://www.alliancegenome.org/disease/DOID:0070301"
                }
            ]
        },
        {
            "disease_id": "DOID:0080046",
            "recommended_name": {
                "id": "DOID:0080046",
                "resource": "Disease Ontology",
                "url": "http://disease-ontology.org/term/DOID:0080046",
                "name": "Stickler syndrome",
                "description": "A syndrome that is characterized by a distinctive facial appearance, eye abnormalities, hearing loss, and joint problems."
            },
            "synonyms": [
                {
                    "id": "MONDO:0007160",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0007160",
                    "name": "Stickler syndrome type 1",
                    "description": ""
                },
                {
                    "id": "MONDO:0007160",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0007160",
                    "name": "Stickler syndrome type 1",
                    "description": ""
                },
                {
                    "id": "MONDO:0007160",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0007160",
                    "name": "STL1",
                    "description": ""
                },
                {
                    "id": "MONDO:0007160",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0007160",
                    "name": "Stickler syndrome, membranous vitreous type",
                    "description": ""
                },
                {
                    "id": "MONDO:0007160",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0007160",
                    "name": "Stickler syndrome, type 1",
                    "description": ""
                },
                {
                    "id": "MONDO:0007160",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0007160",
                    "name": "Stickler syndrome, type I",
                    "description": ""
                },
                {
                    "id": "MONDO:0007160",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0007160",
                    "name": "Stickler syndrome, vitreous type 1",
                    "description": ""
                },
                {
                    "id": "MONDO:0007160",
                    "resource": "MONDO",
                    "url": "https://monarchinitiative.org/disease/MONDO:0007160",
                    "name": "arthroophthalmopathy, hereditary progressive",
                    "description": ""
                },
                {
                    "id": "MIM:108300",
                    "resource": "MIM",
                    "url": "https://www.omim.org/entry/108300",
                    "name": "Stickler syndrome 1",
                    "description": "Stickler syndrome 1"
                }
            ],
            "evidence": [
                {
                    "id": "0080046",
                    "database": "Alliance of Genome Resources",
                    "url": "https://www.alliancegenome.org/disease/DOID:0080046"
                }
            ]
        },
        {
            "disease_id": "DOID:2256",
            "recommended_name": {
                "id": "DOID:2256",
                "resource": "Disease Ontology",
                "url": "http://disease-ontology.org/term/DOID:2256",
                "name": "osteochondrodysplasia",
                "description": "A bone development disease that results_in defective development of cartilage or bone."
            },
            "synonyms": [
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "Cartilage Development disorder",
                    "description": ""
                },
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "Congenital anomaly of cartilage",
                    "description": ""
                },
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "Osteochondrodysplasia syndrome",
                    "description": ""
                },
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "chondrodystrophy",
                    "description": ""
                },
                {
                    "id": "DOID:2256",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:2256",
                    "name": "skeletal dysplasia",
                    "description": ""
                }
            ],
            "evidence": [
                {
                    "id": "2256",
                    "database": "Alliance of Genome Resources",
                    "url": "https://www.alliancegenome.org/disease/DOID:2256"
                }
            ]
        },
        {
            "disease_id": "DOID:8398",
            "recommended_name": {
                "id": "DOID:8398",
                "resource": "Disease Ontology",
                "url": "http://disease-ontology.org/term/DOID:8398",
                "name": "osteoarthritis",
                "description": "An arthritis that has_material_basis_in worn out cartilage located_in joint."
            },
            "synonyms": [
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "Osteoarthrosis and allied disorder",
                    "description": ""
                },
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "degenerative arthritis",
                    "description": ""
                },
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "degenerative joint disease",
                    "description": ""
                },
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "hypertrophic arthritis",
                    "description": ""
                },
                {
                    "id": "DOID:8398",
                    "resource": "Disease Ontology",
                    "url": "http://disease-ontology.org/term/DOID:8398",
                    "name": "osteoarthrosis",
                    "description": ""
                }
            ],
            "evidence": [
                {
                    "id": "8398",
                    "database": "Alliance of Genome Resources",
                    "url": "https://www.alliancegenome.org/disease/DOID:8398"
                }
            ]
        }
    ],
    "snv": [],
    "mutagenesis": [],
    "expression_tissue": [],
    "expression_disease": [],
    "species": [
        {
            "name": "Rattus norvegicus",
            "taxid": 10116,
            "evidence": [
                {
                    "database": "UniProtKB",
                    "id": "P20850",
                    "url": "https://www.uniprot.org/uniprot/P20850"
                }
            ],
            "common_name": "Rat"
        }
    ],
    "gene": [
        {
            "name": "Col9a1",
            "url": "https://www.uniprot.org/uniprot/P20850",
            "locus": {
                "chromosome": "9",
                "start_pos": 26668213,
                "end_pos": 26585034,
                "strand": "-",
                "evidence": [
                    {
                        "database": "Ensembl Gene",
                        "id": "ENSRNOG00000012920",
                        "url": "https://www.ensembl.org/id/ENSRNOG00000012920"
                    }
                ]
            }
        }
    ],
    "crossref": [
        {
            "id": "ENSRNOG00000012920",
            "url": "https://bgee.org/?page=gene&gene_id=ENSRNOG00000012920",
            "database": "Bgee",
            "categories": [
                "Gene Expression"
            ]
        },
        {
            "id": "305104",
            "url": "https://www.ncbi.nlm.nih.gov/gene/305104",
            "database": "GeneID",
            "categories": [
                "Genome Annotation"
            ]
        },
        {
            "id": "IPR001791",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR001791",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR008160",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR008160",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "IPR013320",
            "url": "https://www.ebi.ac.uk/interpro/entry/IPR013320",
            "database": "InterPro",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "rno:305104",
            "url": "https://www.genome.jp/dbget-bin/get_linkdb?-t+pathway+rno:305104",
            "database": "KEGG Pathway",
            "categories": [
                "Enzyme and Pathway"
            ]
        },
        {
            "id": "LFIDCKR",
            "url": "https://omabrowser.org/oma/group/LFIDCKR/",
            "database": "OMA",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "2968414at2759",
            "url": "http://www.orthodb.org/?query=2968414at2759",
            "database": "OrthoDB",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PTHR24023",
            "url": "http://www.pantherdb.org/panther/family.do?clsAccession=PTHR24023",
            "database": "PANTHER",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PTHR24023:SF1082",
            "url": "http://www.pantherdb.org/panther/family.do?clsAccession=PTHR24023:SF1082",
            "database": "PANTHER",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PTHR24637",
            "url": "http://www.pantherdb.org/panther/family.do?clsAccession=PTHR24637",
            "database": "PANTHER",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PTHR24637:SF391",
            "url": "http://www.pantherdb.org/panther/family.do?clsAccession=PTHR24637:SF391",
            "database": "PANTHER",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "PF01391",
            "url": "http://pfam.xfam.org/family/PF01391",
            "database": "Pfam",
            "categories": [
                "Protein Classification"
            ]
        },
        {
            "id": "1309425",
            "url": "http://rgd.mcw.edu/tools/genes/genes_view.cgi?id=1309425",
            "database": "RGD",
            "categories": [
                "Organism-specific"
            ]
        },
        {
            "id": "A0A8I5ZSW9",
            "url": "https://www.uniprot.org/uniprot/A0A8I5ZSW9",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        },
        {
            "id": "A0A8I5ZSW9-1",
            "url": "https://www.uniprot.org/uniprot/A0A8I5ZSW9-1",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        },
        {
            "id": "F1LQ93",
            "url": "https://www.uniprot.org/uniprot/F1LQ93",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        },
        {
            "id": "F1LQ93-1",
            "url": "https://www.uniprot.org/uniprot/F1LQ93-1",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        },
        {
            "id": "P20850",
            "url": "https://www.uniprot.org/uniprot/P20850",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        },
        {
            "id": "P20850-1",
            "url": "https://www.uniprot.org/uniprot/P20850-1",
            "database": "UniProtKB",
            "categories": [
                "Protein"
            ]
        }
    ],
    "sequence": {
        "sequence": "PGQLGNSGKPGQQGPPGEVGPRGPRGLPGSRGPVGPEGSPGIPGKLGPLGSPGLPGLPGPPGLPGMKGDRGVFGEPGPKGEQGASGEEGEAGVRGDLGDMGQPGPKGSVGNPGEPGLRGPEGIRGLPGVEGPRGPPGPRGVQGEQGATGLPGIQGPPGRAPTDQHIKQVCMRVVQEHFAEMAASLKRPDTGASGLPGRPGPPGPPGPPGENGFPGQMGIRGLPGIKGPPGALGLRGPKGDLGEKGERGPPGRGPKGLPGAIGLPGDPGPASYGKNGRDGEQGPPGVAGIPGVPGPPGPPGPPGFCEPASCTLQAGQRAFSKGPDK",
        "length": 325,
        "header": "sp|P20850-1|CO9A1_RAT Isoform 1 of Collagen alpha-1(IX) chain  OS=Rattus norvegicus OX=10116 GN=Col9a1 PE=3 SV=1 CANONICAL=P20850-1"
    },
    "isoforms": [
        {
            "isoform_ac": "A0A8I5ZSW9-1",
            "url": "",
            "sequence": {
                "sequence": "MAWTAWGRGALGLSLMLSGLRLCAAQRGPPGEQGPPGPPGPPGVPGIDGIDGDRGPKGPPGPPGPPGDPGKPGAPGKPGTPGADGLTGPDGSPGSVGPRGQKGEPGVPGSRGFPGRGIPGPPGPPGTTGLPGELGRVGPAGDPGKRGPPGPPGPPGPSGTIGFHDGDPLCPNSCPPGRSGYPGLPGMRGHKGAKGEIGEPGRQGHKGEEGDQGELGEVGAQGPPGAQGLRGITGIVGDKGEKGARGFDGDPGPQGVPGAAGDQGQRGPPGETGPKGDRGIQGSRGIPGSPGPKGDTGLPGVDGRDGIPGMPGTKGEAGKPGPPGDAGLQGLPGVPGIPGAKGVAGEKGNTGPPGKPGQLGSSGKPGQQGPPGEVGPRGPRGLPGSRGPVGPEGSPGIPGKLGPLGSPGLPGLPGPPGLPGMKGDRGVFGEPGPKGEQGASGEEGEAGVRGDLGDMGQPGPKGSVGNPGEPGLRGPEGIRGLPGVEGPRGPPGPRGVQGEQGATGLPGIQGPPGRAPTDQHIKQVCMRVVQEHFAEMAASLKRPDTGASGLPGRPGPPGPPGPPGENGFPGQMGIRGLPGIKGPPGALGLRGPKGDLGEKGERGPPGRGPKGLPGAIGLPGDPGPASYGKNGRDGEQGPPGVAGIPGVPGPPGPPGPPGFCEPASCTLQAGQRAFSKGPDK",
                "length": 680,
                "header": "tr|A0A8I5ZSW9-1|A0A8I5ZSW9_RAT Computationally mapped isoform of Collagen type IX alpha 1 chain  OS=Rattus norvegicus OX=10116 GN=Col9a1 PE=4 SV=1 CANONICAL=P20850-1"
            },
            "locus": {
                "chromosome": "9",
                "start_pos": 26667417,
                "end_pos": 26605363,
                "strand": "-",
                "evidence": [
                    {
                        "database": "Ensembl Transcript",
                        "id": "ENSRNOT00000115769.1",
                        "url": "https://www.ensembl.org/id/ENSRNOT00000115769.1"
                    },
                    {
                        "database": "Ensembl Peptide",
                        "id": "ENSRNOP00000081771.1",
                        "url": "https://www.ensembl.org/id/ENSRNOP00000081771.1"
                    }
                ]
            }
        },
        {
            "isoform_ac": "F1LQ93-1",
            "url": "",
            "sequence": {
                "sequence": "MVTLINPREPTGKMKNFWKIPLFFFACSFLGPCVSATPKRRARFPANSVSHGGSELCPKIRIGQDDLPGFDLISQFQIEKAASRRTIQRVVGSTALQVAYKLGSNVDFRIPTRHLYPSGLPEEYSFLTTFRMTGSTLEKQWNIWQIQDSSGREQVGVKINGQTKSVAFSYKGLDGSLQTAAFLNLQSLFDSRWHKLMIGVERTRATLFIDCIQIDSLPIKPRAQIDADGFAVLGKLVDNPQVSVPFELQWMLIHCDPLRPRRETCHELPVRITTSQTTDERGPPGEQGPPGPPGPPGVPGIDGIDGDRGPKGPPGPPGPPGDPGKPGAPGKPGTPGADGLTGPDGSPGSVGPRGQKGEPGVPGSRGFPGRGIPGPPGPPGTTGLPGELGRVGPAGDPGKRGPPGPPGPPGPSGTIGFHDGDPLCPNSCPPGRSGYPGLPGMRGHKGAKGEIGEPGRQGHKGEEGDQGELGEVGAQGPPGAQGLRGITGIVGDKGEKGARGFDGDPGPQGVPGAAGDQGQRGPPGETGPKGDRGIQGSRGIPGSPGPKGDTGLPGVDGRDGIPGMPGTKGEAGKPGPPGDAGLQGLPGVPGIPGAKGVAGEKGNTGPPGKPGQLGSSGKPGQQGPPGEVGPRGPRGLPGSRGPVGPEGSPGIPGKLGPLGSPGLPGLPGPPGLPGMKGDRGVFGEPGPKGEQGASGEEGEAGVRGDLGDMGQPGPKGSVGNPGEPGLRGPEGIRGLPGVEGPRGPPGPRGVQGEQGATGLPGIQGPPGRAPTDQHIKQVCMRVVQEHFAEMAASLKRPDTGASGLPGRPGPPGPPGPPGENGFPGQMGIRGLPGIKGPPGALGLRGPKGDLGEKGERGPPGRGPKGLPGAIGLPGDPGPASYGKNGRDGEQGPPGVAGIPGVPGPPGPPGPPGFCEPASCTLQAGQRAFSKGPDK",
                "length": 934,
                "header": "tr|F1LQ93-1|F1LQ93_RAT Computationally mapped isoform of Collagen type IX alpha 1 chain  OS=Rattus norvegicus OX=10116 GN=Col9a1 PE=4 SV=3 CANONICAL=P20850-1"
            },
            "locus": {
                "chromosome": "9",
                "start_pos": 26667417,
                "end_pos": 26585240,
                "strand": "-",
                "evidence": [
                    {
                        "database": "Ensembl Transcript",
                        "id": "ENSRNOT00000018116.7",
                        "url": "https://www.ensembl.org/id/ENSRNOT00000018116.7"
                    },
                    {
                        "database": "Ensembl Peptide",
                        "id": "ENSRNOP00000018116.6",
                        "url": "https://www.ensembl.org/id/ENSRNOP00000018116.6"
                    }
                ]
            }
        },
        {
            "isoform_ac": "P20850-1",
            "url": "",
            "sequence": {
                "sequence": "PGQLGNSGKPGQQGPPGEVGPRGPRGLPGSRGPVGPEGSPGIPGKLGPLGSPGLPGLPGPPGLPGMKGDRGVFGEPGPKGEQGASGEEGEAGVRGDLGDMGQPGPKGSVGNPGEPGLRGPEGIRGLPGVEGPRGPPGPRGVQGEQGATGLPGIQGPPGRAPTDQHIKQVCMRVVQEHFAEMAASLKRPDTGASGLPGRPGPPGPPGPPGENGFPGQMGIRGLPGIKGPPGALGLRGPKGDLGEKGERGPPGRGPKGLPGAIGLPGDPGPASYGKNGRDGEQGPPGVAGIPGVPGPPGPPGPPGFCEPASCTLQAGQRAFSKGPDK",
                "length": 325,
                "header": "sp|P20850-1|CO9A1_RAT Isoform 1 of Collagen alpha-1(IX) chain  OS=Rattus norvegicus OX=10116 GN=Col9a1 PE=3 SV=1 CANONICAL=P20850-1"
            },
            "locus": {}
        }
    ],
    "glycosylation": [],
    "phosphorylation": [],
    "glycation": [],
    "keywords": [],
    "publication": [
        {
            "title": "The localization of proteins encoded by CRYM, KIAA1199, UBA52, COL9A3, and COL9A1, genes highly expressed in the cochlea.",
            "journal": "Neuroscience",
            "date": "2008",
            "authors": "Usami S, Takumi Y, Suzuki N, Oguchi T, Oshima A, Suzuki H, Kitoh R, Abe S, Sasaki A, Matsubara A",
            "evidence": [
                {
                    "database": "RefSeq",
                    "id": "NP_001094312",
                    "url": "https://www.ncbi.nlm.nih.gov/protein/NP_001094312"
                }
            ],
            "reference": [
                {
                    "type": "PubMed",
                    "id": "18448257",
                    "url": "https://glygen.org/publication/PubMed/18448257"
                }
            ]
        },
        {
            "title": "Molecular cloning of rat and human type IX collagen cDNA and localization of the alpha 1(IX) gene on the human chromosome 6.",
            "journal": "Eur. J. Biochem.",
            "date": "1989",
            "authors": "Mattei M.-G., Olsen B.R., Kimura T., Stevens J.W., Goldring M.B., Ninomiya Y.",
            "evidence": [
                {
                    "database": "UniProtKB",
                    "id": "P20850",
                    "url": "https://www.uniprot.org/uniprot/P20850"
                }
            ],
            "reference": [
                {
                    "type": "PubMed",
                    "id": "2465149",
                    "url": "https://glygen.org/publication/PubMed/2465149"
                }
            ]
        }
    ],
    "pathway": [
        {
            "id": "rno:305104",
            "name": "",
            "url": "https://www.genome.jp/dbget-bin/get_linkdb?-t+pathway+rno:305104",
            "resource": "KEGG Pathway"
        }
    ],
    "orthologs": [
        {
            "uniprot_canonical_ac": "P20849-1",
            "protein_name": "Collagen alpha-1(IX) chain",
            "gene_name": "COL9A1",
            "tax_id": 9606,
            "organism": "Homo sapiens",
            "sequence": {
                "sequence": "MKTCWKIPVFFFVCSFLEPWASAAVKRRPRFPVNSNSNGGNELCPKIRIGQDDLPGFDLISQFQVDKAASRRAIQRVVGSATLQVAYKLGNNVDFRIPTRNLYPSGLPEEYSFLTTFRMTGSTLKKNWNIWQIQDSSGKEQVGIKINGQTQSVVFSYKGLDGSLQTAAFSNLSSLFDSQWHKIMIGVERSSATLFVDCNRIESLPIKPRGPIDIDGFAVLGKLADNPQVSVPFELQWMLIHCDPLRPRRETCHELPARITPSQTTDERGPPGEQGPPGPPGPPGVPGIDGIDGDRGPKGPPGPPGPAGEPGKPGAPGKPGTPGADGLTGPDGSPGSIGSKGQKGEPGVPGSRGFPGRGIPGPPGPPGTAGLPGELGRVGPVGDPGRRGPPGPPGPPGPRGTIGFHDGDPLCPNACPPGRSGYPGLPGMRGHKGAKGEIGEPGRQGHKGEEGDQGELGEVGAQGPPGAQGLRGITGIVGDKGEKGARGLDGEPGPQGLPGAPGDQGQRGPPGEAGPKGDRGAEGARGIPGLPGPKGDTGLPGVDGRDGIPGMPGTKGEPGKPGPPGDAGLQGLPGVPGIPGAKGVAGEKGSTGAPGKPGQMGNSGKPGQQGPPGEVGPRGPQGLPGSRGELGPVGSPGLPGKLGSLGSPGLPGLPGPPGLPGMKGDRGVVGEPGPKGEQGASGEEGEAGERGELGDIGLPGPKGSAGNPGEPGLRGPEGSRGLPGVEGPRGPPGPRGVQGEQGATGLPGVQGPPGRAPTDQHIKQVCMRVIQEHFAEMAASLKRPDSGATGLPGRPGPPGPPGPPGENGFPGQMGIRGLPGIKGPPGALGLRGPKGDLGEKGERGPPGRGPNGLPGAIGLPGDPGPASYGRNGRDGERGPPGVAGIPGVPGPPGPPGLPGFCEPASCTMQAGQRAFNKGPDP",
                "length": 921
            },
            "evidence": [
                {
                    "id": "45236947",
                    "url": "http://www.informatics.jax.org/homology/45236947",
                    "database": "MGI"
                }
            ],
            "common_name": "Human"
        },
        {
            "uniprot_canonical_ac": "Q05722-1",
            "protein_name": "Collagen alpha-1(IX) chain",
            "gene_name": "Col9a1",
            "tax_id": 10090,
            "organism": "Mus musculus",
            "sequence": {
                "sequence": "MKNFWKISVFFCVCSCLGPWVSATLKRRARFPANSISNGGSELCPKIRIGQDDLPGFDLISQFQIEKAASRRTIQRVVGSTALQVAYKLGSNVDFRIPTRHLYPSGLPEEYSFLTTFRMTGSTLEKHWNIWQIQDSAGREQVGVKINGQTKSVAFSYKGLDGSLQTAAFLNLPSLFDSRWHKLMIGVERTSATLFIDCIRIESLPIKPRGQIDADGFAVLGKLVDNPQVSVPFELQWMLIHCDPLRPRRETCHELPIRITTSQTTDERGPPGEQGPPGPPGPPGVPGIDGIDGDRGPKGPPGPPGPPGDPGKPGAPGKPGTPGADGLTGPDGSPGSVGPRGQKGEPGVPGSRGFPGRGIPGPPGPPGTTGLPGELGRVGPIGDPGKRGPPGPPGPPGPSGTIGFHDGDPLCPNSCPPGRSGYPGLPGMRGHKGAKGEIGEPGRQGHKGEEGDQGELGEVGAQGPPGPQGLRGITGIVGDKGEKGARGFDGEPGPQGIPGAAGDQGQRGPPGETGPKGDRGIQGSRGIPGSPGPKGDTGLPGVDGRDGIPGMPGTKGEAGKPGPPGDVGLQGLPGVPGIPGAKGVAGEKGNTGAPGKPGQLGSSGKPGQQGPPGEVGPRGPRGLPGSRGPVGPEGSPGIPGKLGSVGSPGLPGLPGPPGLPGMKGDRGVFGEPGPKGEQGASGEEGEAGARGDLGDMGQPGPKGSVGNPGEPGLRGPEGIRGLPGVEGPRGPPGPRGMQGEQGATGLPGIQGPPGRAPTDQHIKQVCMRVVQEHFVEMAASLKRPDTGASGLPGRPGPPGPPGPPGENGFPGQMGIRGLPGIKGPPGALGLRGPKGDLGEKGERGPPGRGPKGLPGAIGLPGDPGPASYGKNGRDGEQGPPGVAGIPGVPGPPGPPGPPGFCEPASCTLQSGQRAFSKGPDK",
                "length": 921
            },
            "evidence": [
                {
                    "id": "45236947",
                    "url": "http://www.informatics.jax.org/homology/45236947",
                    "database": "MGI"
                }
            ],
            "common_name": "Mouse"
        }
    ],
    "go_annotation": {
        "categories": [
            {
                "name": "Cellular Component",
                "total": 2,
                "go_terms": [
                    {
                        "name": "extracellular region",
                        "id": "GO:0005576",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0005576",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "collagen type IX trimer",
                        "id": "GO:0005594",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0005594",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    }
                ]
            },
            {
                "name": "Molecular Function",
                "total": 3,
                "go_terms": [
                    {
                        "name": "protein homodimerization activity",
                        "id": "GO:0042803",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0042803",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "metal ion binding",
                        "id": "GO:0046872",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0046872",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "carbohydrate binding",
                        "id": "GO:0030246",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0030246",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    }
                ]
            },
            {
                "name": "Biological Process",
                "total": 7,
                "go_terms": [
                    {
                        "name": "tissue homeostasis",
                        "id": "GO:0001894",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0001894",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "skeletal system development",
                        "id": "GO:0001501",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0001501",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "bone morphogenesis",
                        "id": "GO:0060349",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0060349",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "chondrocyte differentiation",
                        "id": "GO:0002062",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0002062",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "cartilage development",
                        "id": "GO:0051216",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0051216",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "chondrocyte proliferation",
                        "id": "GO:0035988",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0035988",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    },
                    {
                        "name": "growth plate cartilage development",
                        "id": "GO:0003417",
                        "url": "https://www.ebi.ac.uk/QuickGO/term/GO:0003417",
                        "evidence": [
                            {
                                "database": "UniProtKB",
                                "id": "P20850",
                                "url": "https://www.uniprot.org/uniprot/P20850"
                            }
                        ],
                        "pmid": ""
                    }
                ]
            }
        ]
    },
    "pro_annotation": [],
    "site_annotation": [],
    "ptm_annotation": [],
    "interactions": [],
    "reactions": [],
    "reaction_participants": [],
    "reaction_enzymes": [],
    "biomarkers": [],
    "synthesized_glycans": [],
    "cluster_types": [
        {
            "name": "isoformset.uniprotkb",
            "label": "Isoforms"
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        {
            "name": "homologset.mgi",
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