{
    "glytoucan_ac": "G09169SF",
    "number_monosaccharides": 17,
    "number_monosaccharides_suffix": "",
    "mass": 3003.07,
    "mass_pme": 3731.89,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "no",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "",
    "missing_score": 57,
    "glycan_type": "Saccharide",
    "iupac": "alpha-D-Galp-(1->3)-beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->2)[beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->4)]-alpha-D-Manp-(1->3)[alpha-D-Galp-(1->3)-beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->2)[alpha-D-Galp-(1->3)-beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->6)]-alpha-D-Manp-(1->6)]-beta-D-Manp-(1->4)-beta-D-GlcpNAc-(1->4)[alpha-L-Fucp-(1->6)]-?-D-GlcpNAc-(1->",
    "wurcs": "WURCS=2.0/7,17,16/[a2122h-1x_1-5_2*NCC/3=O][a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][a2112h-1a_1-5][a1221m-1a_1-5]/1-2-3-4-2-5-6-2-5-4-2-5-6-2-5-6-7/a4-b1_a6-q1_b4-c1_c3-d1_c6-j1_d2-e1_d4-h1_e4-f1_f3-g1_h4-i1_j2-k1_j6-n1_k4-l1_l3-m1_n4-o1_o3-p1",
    "glycoct": "RES 1b:x-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dgal-HEX-1:5 11b:b-dglc-HEX-1:5 12s:n-acetyl 13b:b-dgal-HEX-1:5 14b:a-dman-HEX-1:5 15b:b-dglc-HEX-1:5 16s:n-acetyl 17b:b-dgal-HEX-1:5 18b:a-dgal-HEX-1:5 19b:b-dglc-HEX-1:5 20s:n-acetyl 21b:b-dgal-HEX-1:5 22b:a-dgal-HEX-1:5 23b:a-lgal-HEX-1:5|6:d LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:9o(3+1)10d 10:6o(4+1)11d 11:11d(2+1)12n 12:11o(4+1)13d 13:5o(6+1)14d 14:14o(2+1)15d 15:15d(2+1)16n 16:15o(4+1)17d 17:17o(3+1)18d 18:14o(6+1)19d 19:19d(2+1)20n 20:19o(4+1)21d 21:21o(3+1)22d 22:1o(6+1)23d",
    "inchi": "InChI=1S/C114H190N6O85/c1-24-53(140)69(156)75(162)104(176-24)174-23-46-91(63(150)47(98(172)177-46)115-25(2)134)194-101-50(118-28(5)137)66(153)89(40(17-130)187-101)199-112-84(171)95(203-114-97(80(167)90(43(20-133)190-114)193-100-49(117-27(4)136)65(152)86(39(16-129)186-100)195-105-76(163)70(157)54(141)31(8-121)178-105)205-103-52(120-30(7)139)68(155)88(42(19-132)189-103)198-111-83(170)94(61(148)37(14-127)184-111)202-108-79(166)73(160)57(144)34(11-124)181-108)62(149)45(191-112)22-175-113-96(204-102-51(119-29(6)138)67(154)87(41(18-131)188-102)197-110-82(169)93(60(147)36(13-126)183-110)201-107-78(165)72(159)56(143)33(10-123)180-107)74(161)58(145)44(192-113)21-173-99-48(116-26(3)135)64(151)85(38(15-128)185-99)196-109-81(168)92(59(146)35(12-125)182-109)200-106-77(164)71(158)55(142)32(9-122)179-106/h24,31-114,121-133,140-172H,8-23H2,1-7H3,(H,115,134)(H,116,135)(H,117,136)(H,118,137)(H,119,138)(H,120,139)/t24-,31+,32+,33+,34+,35+,36+,37+,38+,39+,40+,41+,42+,43+,44+,45+,46+,47+,48+,49+,50+,51+,52+,53+,54-,55-,56-,57-,58+,59-,60-,61-,62+,63+,64+,65+,66+,67+,68+,69+,70-,71-,72-,73-,74-,75-,76+,77+,78+,79+,80-,81+,82+,83+,84-,85+,86+,87+,88+,89+,90+,91+,92-,93-,94-,95-,96-,97-,98?,99+,100-,101-,102-,103-,104+,105-,106+,107+,108+,109-,110-,111-,112-,113-,114+/m0/s1",
    "glycam": "",
    "smiles_isomeric": "C[C@H]1[C@H]([C@H]([C@@H]([C@@H](O1)OC[C@@H]2[C@H]([C@@H]([C@H](C(O2)O)NC(=O)C)O)O[C@H]3[C@@H]([C@H]([C@@H]([C@H](O3)CO)O[C@H]4[C@H]([C@H]([C@@H]([C@H](O4)CO[C@@H]5[C@H]([C@H]([C@@H]([C@H](O5)CO[C@H]6[C@@H]([C@H]([C@@H]([C@H](O6)CO)O[C@H]7[C@@H]([C@H]([C@H]([C@H](O7)CO)O)O[C@@H]8[C@@H]([C@H]([C@H]([C@H](O8)CO)O)O)O)O)O)NC(=O)C)O)O)O[C@H]9[C@@H]([C@H]([C@@H]([C@H](O9)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O[C@@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O)O)O)O)NC(=O)C)O)O[C@@H]1[C@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O)O)O)NC(=O)C)O)O[C@H]1[C@@H]([C@H]([C@@H]([C@H](O1)CO)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O[C@@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O)O)O)O)NC(=O)C)O)O)NC(=O)C)O)O)O",
    "inchi_key": {
        "key": "BEMDAHDLSHQGEP-PEUAVCGHSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/BEMDAHDLSHQGEP-PEUAVCGHSA-N"
    },
    "byonic": "HexNAc(6)Hex(10)dHex(1) % 3003.0729442",
    "gwb": "freeEnd--1?1D-GlcNAc,p(--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p(--2b1D-GlcNAc,p--4b1D-Gal,p--3a1D-Gal,p)--4b1D-GlcNAc,p--4b1D-Gal,p)--6a1D-Man,p(--2b1D-GlcNAc,p--4b1D-Gal,p--3a1D-Gal,p)--6b1D-GlcNAc,p--4b1D-Gal,p--3a1D-Gal,p)--6a1L-Fuc,p$MONO,Und,0,0,freeEnd",
    "species": [],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": "https://glycomotif.glyomics.org/glycomotif/N-glycan"
            },
            "subtype": {
                "name": "Complex",
                "url": "https://glycomotif.glyomics.org/glycomotif/complex_N-glycan"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": "https://glycomotif.glyomics.org/glycomotif/N-glycan"
            },
            "subtype": {
                "name": "Core-fucosylated"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": "https://glycomotif.glyomics.org/glycomotif/N-glycan"
            },
            "subtype": {
                "name": "Tetraantennary"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BYC5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-(1,6)-fucosyltransferase",
            "gene": "FUT8",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BYC5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9WTS2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-(1,6)-fucosyltransferase",
            "gene": "Fut8",
            "gene_link": "https://www.uniprot.org/uniprot/Q9WTS2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10469-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q10469",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921V5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat2",
            "gene_link": "https://www.uniprot.org/uniprot/Q921V5",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P16442-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Histo-blood group ABO system transferase",
            "gene": "ABO",
            "gene_link": "https://www.uniprot.org/uniprot/P16442",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P23336-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "N-acetyllactosaminide alpha-1,3-galactosyltransferase",
            "gene": "Ggta1",
            "gene_link": "https://www.uniprot.org/uniprot/P23336",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P38649-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Histo-blood group ABO system transferase",
            "gene": "Abo",
            "gene_link": "https://www.uniprot.org/uniprot/P38649",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q3V1N9-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-1,3-galactosyltransferase 2",
            "gene": "A3galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q3V1N9",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q4G0N0-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Inactive N-acetyllactosaminide alpha-1,3-galactosyltransferase",
            "gene": "GGTA1",
            "gene_link": "https://www.uniprot.org/uniprot/Q4G0N0",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "U3KPV4-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-1,3-galactosyltransferase 2",
            "gene": "A3GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/U3KPV4",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q812F8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G09169SF",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase B",
            "gene": "Mgat4b",
            "gene_link": "https://www.uniprot.org/uniprot/Q812F8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G09169SF",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G09169SF"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q812G0-1",
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