{
    "glytoucan_ac": "G08569VN",
    "number_monosaccharides": 7,
    "number_monosaccharides_suffix": "",
    "mass": 1572.48,
    "mass_pme": 1936.88,
    "tool_support": {
        "gnome_glygen_oglycans": "yes",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "yes",
        "pdb": "no",
        "gnome_glygen_nglycans": "no",
        "gnome_glycotree_nglycans": "no"
    },
    "fully_determined": "",
    "missing_score": 140,
    "glycan_type": "Saccharide",
    "iupac": "alpha-D-Neup5Ac-(2->3)-beta-D-Galp6S-(1->3)-beta-D-Galp-(1->3)[alpha-D-Neup5Ac-(2->3)-beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->6)]-?-D-GalpNAc-(1->",
    "wurcs": "WURCS=2.0/5,7,6/[a2112h-1x_1-5_2*NCC/3=O][a2112h-1b_1-5][a2112h-1b_1-5_6*OSO/3=O/3=O][Aad21122h-2a_2-6_5*NCC/3=O][a2122h-1b_1-5_2*NCC/3=O]/1-2-3-4-5-2-4/a3-b1_a6-e1_b3-c1_c3-d2_e4-f1_f3-g2",
    "glycoct": "RES 1b:x-dgal-HEX-1:5 2s:n-acetyl 3b:b-dgal-HEX-1:5 4b:b-dgal-HEX-1:5 5b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 6s:n-acetyl 7s:sulfate 8b:b-dglc-HEX-1:5 9s:n-acetyl 10b:b-dgal-HEX-1:5 11b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 12s:n-acetyl LIN 1:1d(2+1)2n 2:1o(3+1)3d 3:3o(3+1)4d 4:4o(3+2)5d 5:5d(5+1)6n 6:4o(6+1)7n 7:1o(6+1)8d 8:8d(2+1)9n 9:8o(4+1)10d 10:10o(3+2)11d 11:11d(5+1)12n",
    "inchi": "InChI=1S/C56H92N4O45S/c1-14(66)57-27-18(70)5-55(53(85)86,102-43(27)31(74)20(72)7-61)104-46-34(77)23(10-64)96-51(39(46)82)99-41-24(11-65)97-49(29(37(41)80)59-16(3)68)92-12-25-35(78)42(30(48(84)94-25)60-17(4)69)100-50-38(81)45(33(76)22(9-63)95-50)101-52-40(83)47(36(79)26(98-52)13-93-106(89,90)91)105-56(54(87)88)6-19(71)28(58-15(2)67)44(103-56)32(75)21(73)8-62/h18-52,61-65,70-84H,5-13H2,1-4H3,(H,57,66)(H,58,67)(H,59,68)(H,60,69)(H,85,86)(H,87,88)(H,89,90,91)/t18-,19-,20+,21+,22+,23+,24+,25+,26+,27+,28+,29+,30+,31+,32+,33-,34-,35-,36-,37+,38+,39+,40+,41+,42+,43+,44+,45-,46-,47-,48?,49+,50-,51-,52-,55-,56-/m0/s1",
    "glycam": "",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H](C[C@@](O[C@H]1[C@@H]([C@@H](CO)O)O)(C(=O)O)O[C@H]2[C@H]([C@H](O[C@H]([C@@H]2O)O[C@@H]3[C@H](O[C@H]([C@@H]([C@H]3O)NC(=O)C)OC[C@@H]4[C@@H]([C@@H]([C@H](C(O4)O)NC(=O)C)O[C@H]5[C@@H]([C@H]([C@H]([C@H](O5)CO)O)O[C@H]6[C@@H]([C@H]([C@H]([C@H](O6)COS(=O)(=O)O)O)O[C@@]7(C[C@@H]([C@H]([C@@H](O7)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)O)CO)CO)O)O",
    "inchi_key": {
        "key": "CQDQSXYZBOSHJQ-ZXOCNWJZSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/CQDQSXYZBOSHJQ-ZXOCNWJZSA-N"
    },
    "byonic": "HexNAc(2)Hex(3)NeuAc(2)Sulpho(1) % 1572.47542827",
    "gwb": "freeEnd--1?1D-GalNAc,p(--3b1D-Gal,p--3b1D-Gal,p(--6S)--3a2D-NeuAc,p)--6b1D-GlcNAc,p--4b1D-Gal,p--3a2D-NeuAc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G08569VN",
                    "url": "https://glycosmos.org/glycans/show/G08569VN"
                }
            ],
            "common_name": "Human"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "O-linked",
                "url": "https://glycomotif.glyomics.org/glycomotif/O-glycan"
            },
            "subtype": {
                "name": "Core 2",
                "url": "https://glycomotif.glyomics.org/glycomotif/core-2_O-glycan"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q96EU7-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "C1GALT1-specific chaperone 1",
            "gene": "C1GALT1C1",
            "gene_link": "https://www.uniprot.org/uniprot/Q96EU7",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9JJ06-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Glycoprotein-N-acetylgalactosamine 3-beta-galactosyltransferase 1",
            "gene": "C1galt1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9JJ06",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9JMG2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "C1GALT1-specific chaperone 1",
            "gene": "C1galt1c1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9JMG2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NS00-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Glycoprotein-N-acetylgalactosamine 3-beta-galactosyltransferase 1",
            "gene": "C1GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NS00",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O95395-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,3-galactosyl-O-glycosyl-glycoprotein beta-1,6-N-acetylglucosaminyltransferase 3",
            "gene": "GCNT3",
            "gene_link": "https://www.uniprot.org/uniprot/O95395",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P97402-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "N-acetyllactosaminide beta-1,6-N-acetylglucosaminyl-transferase",
            "gene": "Gcnt2",
            "gene_link": "https://www.uniprot.org/uniprot/P97402",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q02742-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,3-galactosyl-O-glycosyl-glycoprotein beta-1,6-N-acetylglucosaminyltransferase",
            "gene": "GCNT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q02742",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q09324-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,3-galactosyl-O-glycosyl-glycoprotein beta-1,6-N-acetylglucosaminyltransferase",
            "gene": "Gcnt1",
            "gene_link": "https://www.uniprot.org/uniprot/Q09324",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q5JCT0-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,3-galactosyl-O-glycosyl-glycoprotein beta-1,6-N-acetylglucosaminyltransferase 3",
            "gene": "Gcnt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q5JCT0",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8N0V5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "N-acetyllactosaminide beta-1,6-N-acetylglucosaminyl-transferase",
            "gene": "GCNT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8N0V5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O43286-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 5",
            "gene": "B4GALT5",
            "gene_link": "https://www.uniprot.org/uniprot/O43286",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprot/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60513-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 4",
            "gene": "B4GALT4",
            "gene_link": "https://www.uniprot.org/uniprot/O60513",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprot/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprot/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9JJ04-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 4",
            "gene": "B4galt4",
            "gene_link": "https://www.uniprot.org/uniprot/Q9JJ04",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9JMK0-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 5",
            "gene": "B4galt5",
            "gene_link": "https://www.uniprot.org/uniprot/Q9JMK0",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O88829-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "Lactosylceramide alpha-2,3-sialyltransferase",
            "gene": "St3gal5",
            "gene_link": "https://www.uniprot.org/uniprot/O88829",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P54751-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 1",
            "gene": "St3gal1",
            "gene_link": "https://www.uniprot.org/uniprot/P54751",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P97325-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "CMP-N-acetylneuraminate-beta-1,4-galactoside alpha-2,3-sialyltransferase",
            "gene": "St3gal3",
            "gene_link": "https://www.uniprot.org/uniprot/P97325",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q11201-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 1",
            "gene": "ST3GAL1",
            "gene_link": "https://www.uniprot.org/uniprot/Q11201",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q11203-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "CMP-N-acetylneuraminate-beta-1,4-galactoside alpha-2,3-sialyltransferase",
            "gene": "ST3GAL3",
            "gene_link": "https://www.uniprot.org/uniprot/Q11203",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G08569VN",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G08569VN"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q11204-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G08569VN",
            "protein_name": "CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 2",
            "gene": "St3gal2",
            "gene_link": "https://www.uniprot.org/uniprot/Q11204",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
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