{
    "glytoucan_ac": "G00320ZP",
    "number_monosaccharides": 12,
    "number_monosaccharides_suffix": "",
    "mass": 2513.88,
    "mass_pme": 3130.57,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "yes",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "alpha-D-Neup5Ac-(2->3)-beta-D-Galp-(1->3)[alpha-D-Neup5Ac-(2->6)]-beta-D-GlcpNAc-(1->2)-alpha-D-Manp-(1->3)[alpha-D-Neup5Ac-(2->3)-beta-D-Galp-(1->3)-beta-D-GlcpNAc-(1->2)-alpha-D-Manp-(1->6)]-beta-D-Manp-(1->4)-beta-D-GlcpNAc-(1->4)-beta-D-GlcpNAc-(1->",
    "wurcs": "WURCS=2.0/5,12,11/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O]/1-1-2-3-1-4-5-5-3-1-4-5/a4-b1_b4-c1_c3-d1_c6-i1_d2-e1_e3-f1_e6-h2_f3-g2_i2-j1_j3-k1_k3-l2",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 11s:n-acetyl 12b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 13s:n-acetyl 14b:a-dman-HEX-1:5 15b:b-dglc-HEX-1:5 16s:n-acetyl 17b:b-dgal-HEX-1:5 18b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 19s:n-acetyl LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(3+1)9d 9:9o(3+2)10d 10:10d(5+1)11n 11:7o(6+2)12d 12:12d(5+1)13n 13:5o(6+1)14d 14:14o(2+1)15d 15:15d(2+1)16n 16:15o(3+1)17d 17:17o(3+2)18d 18:18d(5+1)19n",
    "inchi": "InChI=1S/C95H155N7O70/c1-23(113)96-45-30(120)8-93(90(144)145,168-73(45)52(126)33(123)11-103)151-22-44-60(134)72(164-87-68(142)78(59(133)40(18-110)155-87)172-95(92(148)149)10-32(122)47(98-25(3)115)75(170-95)54(128)35(125)13-105)51(102-29(7)119)84(159-44)167-80-65(139)56(130)37(15-107)157-89(80)165-76-61(135)43(160-85(66(76)140)162-70-42(20-112)158-82(49(63(70)137)100-27(5)117)161-69-41(19-111)152-81(143)48(62(69)136)99-26(4)116)21-150-88-79(64(138)55(129)36(14-106)156-88)166-83-50(101-28(6)118)71(57(131)38(16-108)153-83)163-86-67(141)77(58(132)39(17-109)154-86)171-94(91(146)147)9-31(121)46(97-24(2)114)74(169-94)53(127)34(124)12-104/h30-89,103-112,120-143H,8-22H2,1-7H3,(H,96,113)(H,97,114)(H,98,115)(H,99,116)(H,100,117)(H,101,118)(H,102,119)(H,144,145)(H,146,147)(H,148,149)/t30-,31-,32-,33+,34+,35+,36+,37+,38+,39+,40+,41+,42+,43+,44+,45+,46+,47+,48+,49+,50+,51+,52+,53+,54+,55+,56+,57+,58-,59-,60+,61+,62+,63+,64-,65-,66-,67+,68+,69+,70+,71+,72+,73+,74+,75+,76-,77-,78-,79-,80-,81+,82-,83-,84-,85-,86-,87-,88-,89+,93+,94-,95-/m0/s1",
    "glycam": "DNeup5Aca2-3DGalpb1-3[DNeup5Aca2-6]DGlcpNAcb1-2DManpa1-3[DNeup5Aca2-3DGalpb1-3DGlcpNAcb1-2DManpa1-6]DManpb1-4DGlcpNAcb1-4DGlcpNAcb1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H](C[C@@](O[C@H]1[C@@H]([C@@H](CO)O)O)(C(=O)O)OC[C@@H]2[C@H]([C@@H]([C@H]([C@@H](O2)O[C@H]3[C@H]([C@@H]([C@H](O[C@@H]3O[C@H]4[C@@H]([C@H](O[C@H]([C@H]4O)O[C@@H]5[C@H](O[C@H]([C@@H]([C@H]5O)NC(=O)C)O[C@@H]6[C@H](O[C@H]([C@@H]([C@H]6O)NC(=O)C)O)CO)CO)CO[C@@H]7[C@H]([C@H]([C@@H]([C@H](O7)CO)O)O)O[C@H]8[C@@H]([C@H]([C@@H]([C@H](O8)CO)O)O[C@H]9[C@@H]([C@H]([C@H]([C@H](O9)CO)O)O[C@@]1(C[C@@H]([C@H]([C@@H](O1)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)NC(=O)C)O)CO)O)O)NC(=O)C)O[C@H]1[C@@H]([C@H]([C@H]([C@H](O1)CO)O)O[C@@]1(C[C@@H]([C@H]([C@@H](O1)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)O",
    "inchi_key": {
        "key": "UKJDMSMNJKJZKM-ZUDAGGCUSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/UKJDMSMNJKJZKM-ZUDAGGCUSA-N"
    },
    "byonic": "HexNAc(4)Hex(5)NeuAc(3) % 2513.87842253",
    "gwb": "freeEnd--1b1D-GlcNAc,p--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p--2b1D-GlcNAc,p(--3b1D-Gal,p--3a2D-NeuAc,p)--6a2D-NeuAc,p)--6a1D-Man,p--2b1D-GlcNAc,p--3b1D-Gal,p--3a2D-NeuAc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 10116,
            "name": "Rattus norvegicus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "2336",
                    "url": "https://glyconnect.expasy.org/browser/structures/2336"
                }
            ],
            "common_name": "Rat"
        },
        {
            "taxid": 10114,
            "name": "Rattus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G00320ZP",
                    "url": "https://glycosmos.org/glycans/show/G00320ZP"
                }
            ],
            "common_name": "Rat"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": "https://glycomotif.glyomics.org/glycomotif/N-glycan"
            },
            "subtype": {
                "name": "Biantennary"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": "https://glycomotif.glyomics.org/glycomotif/N-glycan"
            },
            "subtype": {
                "name": "Complex",
                "url": "https://glycomotif.glyomics.org/glycomotif/complex_N-glycan"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P97325-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "CMP-N-acetylneuraminate-beta-1,4-galactoside alpha-2,3-sialyltransferase",
            "gene": "St3gal3",
            "gene_link": "https://www.uniprot.org/uniprot/P97325",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q11203-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "CMP-N-acetylneuraminate-beta-1,4-galactoside alpha-2,3-sialyltransferase",
            "gene": "ST3GAL3",
            "gene_link": "https://www.uniprot.org/uniprot/Q11203",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q11206-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 4",
            "gene": "ST3GAL4",
            "gene_link": "https://www.uniprot.org/uniprot/Q11206",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8VIB3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Type 2 lactosamine alpha-2,3-sialyltransferase",
            "gene": "St3gal6",
            "gene_link": "https://www.uniprot.org/uniprot/Q8VIB3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91Y74-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 4",
            "gene": "St3gal4",
            "gene_link": "https://www.uniprot.org/uniprot/Q91Y74",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Y274-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Type 2 lactosamine alpha-2,3-sialyltransferase",
            "gene": "ST3GAL6",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Y274",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O43825-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Beta-1,3-galactosyltransferase 2",
            "gene": "B3GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O43825",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O54905-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Beta-1,3-galactosyltransferase 2",
            "gene": "B3galt2",
            "gene_link": "https://www.uniprot.org/uniprot/O54905",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9JI67-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Beta-1,3-galactosyltransferase 5",
            "gene": "B3galt5",
            "gene_link": "https://www.uniprot.org/uniprot/Q9JI67",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Y2C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Beta-1,3-galactosyltransferase 5",
            "gene": "B3GALT5",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Y2C3",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10469-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT2",
            "gene_link": "https://www.uniprot.org/uniprot/Q10469",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921V5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat2",
            "gene_link": "https://www.uniprot.org/uniprot/Q921V5",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "N-acetylglucosaminyldiphosphodolichol N-acetylglucosaminyltransferase",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14 homolog",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14 homolog",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00320ZP",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00320ZP"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00320ZP",
            "protein_name": "Putative bifunctional UDP-N-acetylglucosamine transferase and deubiquitinase ALG13",
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            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
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