{
    "glytoucan_ac": "G10503MP",
    "number_monosaccharides": 7,
    "number_monosaccharides_suffix": "",
    "mass": 1328.37,
    "mass_pme": 1636.71,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "no",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "yes",
        "gnome_glygen_nglycans": "no",
        "gnome_glycotree_nglycans": "no"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "beta-D-GlcpA3S-(1->3)-beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->3)-beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->3)-beta-D-Galp-(1->4)-beta-D-Glcp-(1->",
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    "glycoct": "RES 1b:b-dglc-HEX-1:5 2b:b-dgal-HEX-1:5 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dgal-HEX-1:5 6b:b-dglc-HEX-1:5 7s:n-acetyl 8b:b-dgal-HEX-1:5 9b:b-dglc-HEX-1:5|6:a 10s:sulfate LIN 1:1o(4+1)2d 2:2o(3+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6d(2+1)7n 7:6o(4+1)8d 8:8o(3+1)9d 9:9o(3+1)10n",
    "inchi": "InChI=1S/C46H76N2O40S/c1-9(55)47-17-22(60)31(15(7-53)79-42(17)85-35-20(58)12(4-50)78-45(28(35)66)83-33-14(6-52)75-40(71)25(63)24(33)62)81-43-27(65)34(19(57)11(3-49)76-43)84-41-18(48-10(2)56)23(61)32(16(8-54)80-41)82-44-29(67)36(21(59)13(5-51)77-44)86-46-30(68)37(88-89(72,73)74)26(64)38(87-46)39(69)70/h11-38,40-46,49-54,57-68,71H,3-8H2,1-2H3,(H,47,55)(H,48,56)(H,69,70)(H,72,73,74)/t11-,12-,13-,14-,15-,16-,17-,18-,19+,20+,21+,22-,23-,24-,25-,26+,27-,28-,29-,30-,31-,32-,33-,34+,35+,36+,37+,38+,40-,41+,42+,43+,44+,45+,46-/m1/s1",
    "glycam": "DGlcpA[3S]b1-3DGalpb1-4DGlcpNAcb1-3DGalpb1-4DGlcpNAcb1-3DGalpb1-4DGlcpb1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H]([C@@H]([C@H](O[C@H]1O[C@H]2[C@H]([C@H](O[C@H]([C@@H]2O)O[C@@H]3[C@H](O[C@H]([C@@H]([C@H]3O)O)O)CO)CO)O)CO)O[C@H]4[C@@H]([C@H]([C@H]([C@H](O4)CO)O)O[C@H]5[C@@H]([C@H]([C@@H]([C@H](O5)CO)O[C@H]6[C@@H]([C@H]([C@H]([C@H](O6)CO)O)O[C@H]7[C@@H]([C@H]([C@@H]([C@H](O7)C(=O)O)O)OS(=O)(=O)O)O)O)O)NC(=O)C)O)O",
    "inchi_key": {
        "key": "UMXJZILSAKZPGU-IZHGJYKOSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/UMXJZILSAKZPGU-IZHGJYKOSA-N"
    },
    "byonic": "Hex(4)HexNAc(2)HexA(1)S(1) % 1328.36950656",
    "gwb": "freeEnd--1b1D-Glc,p--4b1D-Gal,p--3b1D-GlcNAc,p--4b1D-Gal,p--3b1D-GlcNAc,p--4b1D-Gal,p--3b1D-GlcA,p--3S$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "Direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G10503MP",
                    "url": "https://glycosmos.org/glycans/show/G10503MP"
                }
            ],
            "common_name": "Human"
        },
        {
            "taxid": 10090,
            "name": "Mus musculus",
            "annotation_category": "Direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G10503MP",
                    "url": "https://glycosmos.org/glycans/show/G10503MP"
                }
            ],
            "common_name": "Mouse"
        },
        {
            "taxid": 10114,
            "name": "Rattus",
            "annotation_category": "Direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G10503MP",
                    "url": "https://glycosmos.org/glycans/show/G10503MP"
                }
            ],
            "common_name": "rats"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "Glycosphingolipid"
            },
            "subtype": {
                "name": "Neo-lacto series"
            }
        }
    ],
    "enzyme": [],
    "crossref": [
        {
            "id": "91856241",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/91856241",
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        {
            "id": "252289219",
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            "database": "PubChem Substance"
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        {
            "id": "G00390",
            "url": "https://www.genome.jp/dbget-bin/www_bget?gl:G00390",
            "database": "KEGG Glycan"
        },
        {
            "id": "5173",
            "url": "http://www.glycosciences.de/database/start.php?action=explore_linucsid&linucsid=5173",
            "database": "Glycosciences.de"
        },
        {
            "id": "11604",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+11604",
            "database": "CarbBank"
        },
        {
            "id": "1250",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+1250",
            "database": "CarbBank"
        },
        {
            "id": "16706",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+16706",
            "database": "CarbBank"
        },
        {
            "id": "17005",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+17005",
            "database": "CarbBank"
        },
        {
            "id": "17018",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+17018",
            "database": "CarbBank"
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        {
            "id": "18170",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+18170",
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        {
            "id": "19222",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+19222",
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        {
            "id": "19225",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+19225",
            "database": "CarbBank"
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        {
            "id": "20332",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+20332",
            "database": "CarbBank"
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        {
            "id": "20343",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+20343",
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        {
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            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+22180",
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        {
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            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+22189",
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            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+28511",
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            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+35287",
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        {
            "id": "35441",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+35441",
            "database": "CarbBank"
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        {
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            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+35985",
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        {
            "id": "37246",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+37246",
            "database": "CarbBank"
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        {
            "id": "37270",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+37270",
            "database": "CarbBank"
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        {
            "id": "37650",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+37650",
            "database": "CarbBank"
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        {
            "id": "38295",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+38295",
            "database": "CarbBank"
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        {
            "id": "42775",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+42775",
            "database": "CarbBank"
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        {
            "id": "48907",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+48907",
            "database": "CarbBank"
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        {
            "id": "147173",
            "url": "https://www.ebi.ac.uk/chebi/searchId.do?chebiId=147173",
            "database": "ChEBI"
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        {
            "id": "G10503MP",
            "url": "https://glytoucan.org/Structures/Glycans/G10503MP",
            "database": "GlyTouCan"
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        {
            "id": "G10503MP",
            "url": "https://gnome.glyomics.org/restrictions/GlyGen.StructureBrowser.html?focus=G10503MP",
            "database": "GNOme"
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        {
            "id": "G10503MP",
            "url": "https://glycosmos.org/glycans/show/G10503MP",
            "database": "GlyCosmos"
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    "publication": [],
    "residues": [],
    "composition": [
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            "name": "6-Deoxy-Hexose",
            "residue": "dhex",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
        },
        {
            "name": "N-Glycolyl-Neuraminic Acid",
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            "cid": "440001",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
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            "cid": "1061",
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            "name": "Pentose",
            "residue": "pent",
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            "cid": "229",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
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            "name": "Sulfate",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
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            "name": "Other",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/17106"
        },
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            "cid": "445724",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/445724"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6036"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/11850"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/165880"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/5793"
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        {
            "name": "D-Glucuronic acid",
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            "cid": "94715",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/94715"
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        {
            "name": "D-Glucosamine",
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            "cid": "439213",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439213"
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            "name": "N-Acetyl-D-Glucosamine",
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            "cid": "439174",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439174"
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        {
            "name": "N-Acetyl-D-glucosaminitol",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/165206"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206"
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            "name": "N-Acetylhexosamine",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
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            "name": "Hexuronic Acid",
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            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
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        {
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739"
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            "name": "Hexitol",
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            "name": "L-Iduronic acid",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/441039"
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            "name": "D-Mannose",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/18950"
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            "name": "D-Mannitol",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6251"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440049"
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        {
            "name": "N-Acetyl-D-mannosamine",
            "residue": "mannac",
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            "cid": "439281",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439281"
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        {
            "name": "Methyl",
            "residue": "me",
            "count": 0
        },
        {
            "name": "N-Acetyl-Neuraminic Acid",
            "residue": "neuac",
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            "cid": "439197",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
        },
        {
            "name": "N-Glycolyl-Neuraminic Acid",
            "residue": "neugc",
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            "cid": "440001",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
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        {
            "name": "Phosphate",
            "residue": "p",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
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        {
            "name": "Pentose",
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            "cid": "229",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
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            "name": "Sulfate",
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            "cid": "1117",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
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        {
            "name": "Sialic acid",
            "residue": "sia",
            "count": 0,
            "cid": "906",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/906"
        },
        {
            "name": "Other",
            "residue": "other",
            "count": 0
        },
        {
            "name": "Floating substituent",
            "residue": "x",
            "count": 0
        },
        {
            "name": "D-Xylose",
            "residue": "xyl",
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            "cid": "135191",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/135191"
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        {
            "name": "Ketodeoxynononic acid",
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            "cid": "13991616",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/13991616"
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    ],
    "motifs": [
        {
            "id": "GGM.000001",
            "name": "Type 2 LN",
            "synonym": "N-acetyllactosamine",
            "keywords": "LacNAc; N-acetyllactosamine; N-acetyllactosamine type 2"
        },
        {
            "id": "GGM.000002",
            "name": "Type 2 LN2",
            "synonym": "i antigen",
            "keywords": "LacNAc; N-acetyllactosamine; N-acetyllactosamine type 2"
        },
        {
            "id": "GGM.000058",
            "name": "HNK-1 antigen",
            "synonym": "",
            "keywords": ""
        }
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    "interactions": [],
    "expression": [],
    "subsumption": [
        {
            "related_accession": "G10667BL",
            "relationship": "ancestor"
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        {
            "related_accession": "G31791WL",
            "relationship": "ancestor"
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            "related_accession": "G43040ZY",
            "relationship": "ancestor"
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            "related_accession": "G53005ZJ",
            "relationship": "ancestor"
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        {
            "related_accession": "G55493MW",
            "relationship": "ancestor"
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        {
            "related_accession": "G63386TM",
            "relationship": "ancestor"
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