{
    "glytoucan_ac": "G10209QU",
    "number_monosaccharides": 8,
    "number_monosaccharides_suffix": "",
    "mass": 1421.52,
    "mass_pme": 1771.91,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "yes",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "yes",
        "gnome_glygen_nglycans": "yes",
        "gnome_glycotree_nglycans": "yes"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "beta-D-Galp-(1->3)-beta-D-GlcpNAc-(1->2)-alpha-D-Manp-(1->3)[alpha-D-Manp-(1->6)]-beta-D-Manp-(1->4)-beta-D-GlcpNAc-(1->4)[alpha-L-Fucp-(1->6)]-beta-D-GlcpNAc-(1->",
    "wurcs": "WURCS=2.0/5,8,7/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][a1221m-1a_1-5]/1-1-2-3-1-4-3-5/a4-b1_a6-h1_b4-c1_c3-d1_c6-g1_d2-e1_e3-f1",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dman-HEX-1:5 11b:a-lgal-HEX-1:5|6:d LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(3+1)9d 9:5o(6+1)10d 10:1o(6+1)11d",
    "inchi": "InChI=1S/C54H91N3O40/c1-12-26(66)34(74)38(78)50(85-12)84-11-22-43(32(72)23(47(82)86-22)55-13(2)63)93-48-24(56-14(3)64)33(73)42(20(9-62)91-48)94-53-41(81)45(31(71)21(92-53)10-83-51-39(79)35(75)27(67)16(5-58)88-51)96-54-46(37(77)29(69)18(7-60)90-54)97-49-25(57-15(4)65)44(30(70)19(8-61)87-49)95-52-40(80)36(76)28(68)17(6-59)89-52/h12,16-54,58-62,66-82H,5-11H2,1-4H3,(H,55,63)(H,56,64)(H,57,65)/t12-,16+,17+,18+,19+,20+,21+,22+,23+,24+,25+,26+,27+,28-,29+,30+,31+,32+,33+,34+,35-,36-,37-,38-,39-,40+,41-,42+,43+,44+,45-,46-,47+,48-,49-,50+,51-,52-,53-,54+/m0/s1",
    "glycam": "DGalpb1-3DGlcpNAcb1-2DManpa1-3[DManpa1-6]DManpb1-4DGlcpNAcb1-4[LFucpa1-6]DGlcpNAcb1-OH",
    "smiles_isomeric": "C[C@H]1[C@H]([C@H]([C@@H]([C@@H](O1)OC[C@@H]2[C@H]([C@@H]([C@H]([C@@H](O2)O)NC(=O)C)O)O[C@H]3[C@@H]([C@H]([C@@H]([C@H](O3)CO)O[C@H]4[C@H]([C@H]([C@@H]([C@H](O4)CO[C@@H]5[C@H]([C@H]([C@@H]([C@H](O5)CO)O)O)O)O)O[C@@H]6[C@H]([C@H]([C@@H]([C@H](O6)CO)O)O)O[C@H]7[C@@H]([C@H]([C@@H]([C@H](O7)CO)O)O[C@H]8[C@@H]([C@H]([C@H]([C@H](O8)CO)O)O)O)NC(=O)C)O)O)NC(=O)C)O)O)O",
    "inchi_key": {
        "key": "JBAWMUVCLDJWEL-ZCHFQZOHSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/JBAWMUVCLDJWEL-ZCHFQZOHSA-N"
    },
    "byonic": "Hex(4)HexNAc(3)dHex(1) % 1421.51788538",
    "gwb": "freeEnd--1b1D-GlcNAc,p(--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p--2b1D-GlcNAc,p--3b1D-Gal,p)--6a1D-Man,p)--6a1L-Fuc,p$MONO,Und,0,0,freeEnd",
    "species": [],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=N-glycan&jump_to_nav=true"
            },
            "subtype": {
                "name": "Core-fucosylated"
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9WTS2-1",
            "protein_name": "Alpha-(1,6)-fucosyltransferase",
            "gene": "Fut8",
            "gene_link": "https://www.uniprot.org/uniprot/Q9WTS2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BYC5-1",
            "protein_name": "Alpha-(1,6)-fucosyltransferase",
            "gene": "FUT8",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BYC5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprot/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprot/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Y2C3-1",
            "protein_name": "Beta-1,3-galactosyltransferase 5",
            "gene": "B3GALT5",
            "gene_link": "https://www.uniprot.org/uniprot/Q9Y2C3",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9JI67-1",
            "protein_name": "Beta-1,3-galactosyltransferase 5",
            "gene": "B3galt5",
            "gene_link": "https://www.uniprot.org/uniprot/Q9JI67",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O54905-1",
            "protein_name": "Beta-1,3-galactosyltransferase 2",
            "gene": "B3galt2",
            "gene_link": "https://www.uniprot.org/uniprot/O54905",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O43825-1",
            "protein_name": "Beta-1,3-galactosyltransferase 2",
            "gene": "B3GALT2",
            "gene_link": "https://www.uniprot.org/uniprot/O43825",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P42867-1",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprot/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "protein_name": "Putative bifunctional UDP-N-acetylglucosamine transferase and deubiquitinase ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "protein_name": "Putative bifunctional UDP-N-acetylglucosamine transferase and deubiquitinase ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14 homolog",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14 homolog",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "protein_name": "N-acetylglucosaminyldiphosphodolichol N-acetylglucosaminyltransferase",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "GLY_000284",
                    "database": "GlyGen",
                    "url": "https://data.glygen.org/GLY_000284"
                }
            ]
        }
    ],
    "crossref": [
        {
            "id": "91857640",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/91857640",
            "database": "PubChem Compound"
        },
        {
            "id": "252290884",
            "url": "https://pubchem.ncbi.nlm.nih.gov/substance/252290884",
            "database": "PubChem Substance"
        },
        {
            "id": "24952",
            "url": "http://www.glycosciences.de/database/start.php?action=explore_linucsid&linucsid=24952",
            "database": "Glycosciences.de"
        },
        {
            "id": "147105",
            "url": "https://www.ebi.ac.uk/chebi/searchId.do?chebiId=147105",
            "database": "ChEBI"
        },
        {
            "id": "G10209QU",
            "url": "https://glytoucan.org/Structures/Glycans/G10209QU",
            "database": "GlyTouCan"
        },
        {
            "id": "G10209QU",
            "url": "https://glygen.ccrc.uga.edu/sandbox/explore.html?G10209QU",
            "database": "SandBox"
        },
        {
            "id": "G10209QU",
            "url": "https://gnome.glyomics.org/restrictions/GlyGen.StructureBrowser.html?focus=G10209QU",
            "database": "GNOme"
        },
        {
            "id": "G10209QU",
            "url": "https://glycosmos.org/glycans/show/G10209QU",
            "database": "GlyCosmos"
        }
    ],
    "glycoprotein": [],
    "publication": [],
    "residues": [
        {
            "id": "N1",
            "name": "N-glycan_a-D-Manp_1",
            "attachedby": "rxn.Q9H553-1",
            "detachedby": "",
            "parentid": "NC"
        },
        {
            "id": "N1",
            "name": "N-glycan_a-D-Manp_1",
            "attachedby": "rxn.Q9DBE8-1",
            "detachedby": "",
            "parentid": "NC"
        },
        {
            "id": "N15",
            "name": "N-glycan_a-L-Fucp_15",
            "attachedby": "rxn.Q9WTS2-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "N15",
            "name": "N-glycan_a-L-Fucp_15",
            "attachedby": "rxn.Q9BYC5-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "N2",
            "name": "N-glycan_b-D-GlcpNAc_2",
            "attachedby": "rxn.P27808-1",
            "detachedby": "",
            "parentid": "N1"
        },
        {
            "id": "N2",
            "name": "N-glycan_b-D-GlcpNAc_2",
            "attachedby": "rxn.P26572-1",
            "detachedby": "",
            "parentid": "N1"
        },
        {
            "id": "N4",
            "name": "N-glycan_a-D-Manp_4",
            "attachedby": "rxn.Q9H553-1",
            "detachedby": "",
            "parentid": "NC"
        },
        {
            "id": "N4",
            "name": "N-glycan_a-D-Manp_4",
            "attachedby": "rxn.Q9DBE8-1",
            "detachedby": "",
            "parentid": "NC"
        },
        {
            "id": "N44",
            "name": "N-glycan_b-D-Galp_44",
            "attachedby": "rxn.Q9Y2C3-1",
            "detachedby": "",
            "parentid": "N2"
        },
        {
            "id": "N44",
            "name": "N-glycan_b-D-Galp_44",
            "attachedby": "rxn.Q9JI67-1",
            "detachedby": "",
            "parentid": "N2"
        },
        {
            "id": "N44",
            "name": "N-glycan_b-D-Galp_44",
            "attachedby": "rxn.O54905-1",
            "detachedby": "",
            "parentid": "N2"
        },
        {
            "id": "N44",
            "name": "N-glycan_b-D-Galp_44",
            "attachedby": "rxn.O43825-1",
            "detachedby": "",
            "parentid": "N2"
        },
        {
            "id": "NA",
            "name": "N-glycan_core_D-GlcNAc_A",
            "attachedby": "rxn.Q9H3H5-1",
            "detachedby": "",
            "parentid": "0"
        },
        {
            "id": "NA",
            "name": "N-glycan_core_D-GlcNAc_A",
            "attachedby": "rxn.P42867-1",
            "detachedby": "",
            "parentid": "0"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q9NP73-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q9D8C3-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q9D081-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q96F25-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q8BML3-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NC",
            "name": "N-glycan_core_b-D-Manp_C",
            "attachedby": "rxn.Q9BT22-1",
            "detachedby": "",
            "parentid": "NB"
        },
        {
            "id": "NC",
            "name": "N-glycan_core_b-D-Manp_C",
            "attachedby": "rxn.Q921Q3-1",
            "detachedby": "",
            "parentid": "NB"
        }
    ],
    "composition": [
        {
            "name": "6-Deoxy-Hexose",
            "residue": "dhex",
            "count": 1,
            "cid": "840",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
        },
        {
            "name": "Hexose",
            "residue": "hex",
            "count": 4,
            "cid": "206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206"
        },
        {
            "name": "N-Acetylhexosamine",
            "residue": "hexnac",
            "count": 3,
            "cid": "899",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
        },
        {
            "name": "Hexuronic Acid",
            "residue": "hexa",
            "count": 0,
            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
        },
        {
            "name": "2-Amino-2-Deoxy-Hexose",
            "residue": "hexn",
            "count": 0,
            "cid": "739",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739"
        },
        {
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
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            "name": "N-Glycolyl-Neuraminic Acid",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
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        {
            "name": "Pentose",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
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            "name": "6-Deoxy-Hexitol",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6036"
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            "name": "D-Glucuronic acid",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/94715"
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        {
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
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        {
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/18950"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6251"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440049"
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        {
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439281"
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        {
            "name": "Methyl",
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
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        {
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
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        {
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/906"
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        {
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/135191"
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        {
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/13991616"
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            "keywords": "LacNAc; N-acetyllactosamine; N-acetyllactosamine type 1"
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}