{
    "glytoucan_ac": "G04690JX",
    "number_monosaccharides": 4,
    "number_monosaccharides_suffix": "",
    "mass": 675.26,
    "mass_pme": 843.45,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "no",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "no",
        "gnome_glygen_nglycans": "no",
        "gnome_glycotree_nglycans": "no"
    },
    "fully_determined": "",
    "missing_score": 492,
    "glycan_type": "Saccharide",
    "iupac": "alpha-L-Fucp-(1->2)-beta-D-Galp-(1->4)[alpha-L-Fucp-(1->3)]-?-D-GlcNAc",
    "wurcs": "WURCS=2.0/3,4,3/[u2122h_2*NCC/3=O][a1221m-1a_1-5][a2112h-1b_1-5]/1-2-3-2/a3-b1_a4-c1_c2-d1",
    "glycoct": "RES 1b:x-dglc-HEX-x:x 2s:n-acetyl 3b:a-lgal-HEX-1:5|6:d 4b:b-dgal-HEX-1:5 5b:a-lgal-HEX-1:5|6:d LIN 1:1d(2+1)2n 2:1o(3+1)3d 3:1o(4+1)4d 4:4o(2+1)5d",
    "inchi": "",
    "glycam": "",
    "smiles_isomeric": "",
    "inchi_key": {},
    "byonic": "Hex(1)HexNAc(1)dHex(2) % 675.258578545",
    "gwb": "freeEnd--?D-GlcNAc(--3a1L-Fuc,p)--4b1D-Gal,p--2a1L-Fuc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "Direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G04690JX",
                    "url": "https://glycosmos.org/glycans/show/G04690JX"
                }
            ],
            "common_name": "Human"
        },
        {
            "taxid": 10090,
            "name": "Mus musculus",
            "annotation_category": "Direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G04690JX",
                    "url": "https://glycosmos.org/glycans/show/G04690JX"
                }
            ],
            "common_name": "Mouse"
        },
        {
            "taxid": 10114,
            "name": "Rattus",
            "annotation_category": "Subsumption",
            "evidence": [
                {
                    "database": "GNOme",
                    "id": "G00052MO",
                    "url": "https://www.glygen.org/glycan/G00052MO"
                }
            ],
            "common_name": "rats"
        },
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "Subsumption",
            "evidence": [
                {
                    "database": "GNOme",
                    "id": "G00052MO",
                    "url": "https://www.glygen.org/glycan/G00052MO"
                }
            ],
            "common_name": "Human"
        },
        {
            "taxid": 10090,
            "name": "Mus musculus",
            "annotation_category": "Subsumption",
            "evidence": [
                {
                    "database": "GNOme",
                    "id": "G00052MO",
                    "url": "https://www.glygen.org/glycan/G00052MO"
                }
            ],
            "common_name": "Mouse"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "Other",
                "url": ""
            },
            "subtype": {
                "name": "Other",
                "url": ""
            }
        }
    ],
    "enzyme": [],
    "crossref": [
        {
            "id": "2611",
            "url": "http://www.glycosciences.de/database/start.php?action=explore_linucsid&linucsid=2611",
            "database": "Glycosciences.de"
        },
        {
            "id": "10641",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+10641",
            "database": "CarbBank"
        },
        {
            "id": "11852",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+11852",
            "database": "CarbBank"
        },
        {
            "id": "14052",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+14052",
            "database": "CarbBank"
        },
        {
            "id": "14396",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+14396",
            "database": "CarbBank"
        },
        {
            "id": "14848",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+14848",
            "database": "CarbBank"
        },
        {
            "id": "14885",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+14885",
            "database": "CarbBank"
        },
        {
            "id": "15543",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+15543",
            "database": "CarbBank"
        },
        {
            "id": "15824",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+15824",
            "database": "CarbBank"
        },
        {
            "id": "29561",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+29561",
            "database": "CarbBank"
        },
        {
            "id": "32811",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+32811",
            "database": "CarbBank"
        },
        {
            "id": "38101",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+38101",
            "database": "CarbBank"
        },
        {
            "id": "4934",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+4934",
            "database": "CarbBank"
        },
        {
            "id": "49591",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+49591",
            "database": "CarbBank"
        },
        {
            "id": "8695",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+8695",
            "database": "CarbBank"
        },
        {
            "id": "8906",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+8906",
            "database": "CarbBank"
        },
        {
            "id": "9917",
            "url": "https://www.genome.jp/dbget-bin/www_bget?carbbank+9917",
            "database": "CarbBank"
        },
        {
            "id": "20538",
            "url": "http://csdb.glycoscience.ru/bacterial/core/search_id.php?mode=record&id_list=20538",
            "database": "BCSDB"
        },
        {
            "id": "23173",
            "url": "http://csdb.glycoscience.ru/bacterial/core/search_id.php?mode=record&id_list=23173",
            "database": "BCSDB"
        },
        {
            "id": "6920",
            "url": "http://csdb.glycoscience.ru/bacterial/core/search_id.php?mode=record&id_list=6920",
            "database": "BCSDB"
        },
        {
            "id": "G04690JX",
            "url": "https://glytoucan.org/Structures/Glycans/G04690JX",
            "database": "GlyTouCan"
        },
        {
            "id": "G04690JX",
            "url": "https://gnome.glyomics.org/restrictions/GlyGen.StructureBrowser.html?focus=G04690JX",
            "database": "GNOme"
        },
        {
            "id": "G04690JX",
            "url": "https://glycosmos.org/glycans/show/G04690JX",
            "database": "GlyCosmos"
        }
    ],
    "glycoprotein": [],
    "publication": [
        {
            "title": "An enzymatic ruler modulates Lewis antigen glycosylation of Helicobacter pylori LPS during persistent infection.",
            "journal": "Proceedings of the National Academy of Sciences of the United States of America",
            "date": "2006",
            "authors": "Nilsson C, Skoglund A, Moran AP, Annuk H, Engstrand L, Normark S",
            "evidence": [
                {
                    "database": "GlyTouCan",
                    "id": "G04690JX",
                    "url": "https://glytoucan.org/Structures/Glycans/G04690JX"
                }
            ],
            "reference": [
                {
                    "type": [
                        "PubMed"
                    ],
                    "id": "16477004",
                    "url": "https://glygen.org/publication/PubMed/16477004"
                }
            ]
        },
        {
            "title": "Relevance of fucosylation and Lewis antigen expression in the bacterial gastroduodenal pathogen Helicobacter pylori.",
            "journal": "Carbohydrate research",
            "date": "2008",
            "authors": "Moran AP",
            "evidence": [
                {
                    "database": "GlyTouCan",
                    "id": "G04690JX",
                    "url": "https://glytoucan.org/Structures/Glycans/G04690JX"
                }
            ],
            "reference": [
                {
                    "type": [
                        "PubMed"
                    ],
                    "id": "18279843",
                    "url": "https://glygen.org/publication/PubMed/18279843"
                }
            ]
        },
        {
            "title": "Cloning and heterologous expression of an alpha1,3-fucosyltransferase gene from the gastric pathogen Helicobacter pylori.",
            "journal": "The Journal of biological chemistry",
            "date": "1997",
            "authors": "Ge Z, Chan NW, Palcic MM, Taylor DE",
            "evidence": [
                {
                    "database": "GlyTouCan",
                    "id": "G04690JX",
                    "url": "https://glytoucan.org/Structures/Glycans/G04690JX"
                }
            ],
            "reference": [
                {
                    "type": [
                        "PubMed"
                    ],
                    "id": "9261149",
                    "url": "https://glygen.org/publication/PubMed/9261149"
                }
            ]
        }
    ],
    "residues": [],
    "composition": [
        {
            "name": "6-Deoxy-Hexose",
            "residue": "dhex",
            "count": 2,
            "cid": "840",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
        },
        {
            "name": "Hexose",
            "residue": "hex",
            "count": 1,
            "cid": "206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206"
        },
        {
            "name": "N-Acetylhexosamine",
            "residue": "hexnac",
            "count": 1,
            "cid": "899",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
        },
        {
            "name": "Hexuronic Acid",
            "residue": "hexa",
            "count": 0,
            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
        },
        {
            "name": "2-Amino-2-Deoxy-Hexose",
            "residue": "hexn",
            "count": 0,
            "cid": "739",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739"
        },
        {
            "name": "N-Acetyl-Neuraminic Acid",
            "residue": "neuac",
            "count": 0,
            "cid": "439197",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
        },
        {
            "name": "N-Glycolyl-Neuraminic Acid",
            "residue": "neugc",
            "count": 0,
            "cid": "440001",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
        },
        {
            "name": "Phosphate",
            "residue": "p",
            "count": 0,
            "cid": "1061",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
        },
        {
            "name": "Pentose",
            "residue": "pent",
            "count": 0,
            "cid": "229",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
        },
        {
            "name": "Sulfate",
            "residue": "s",
            "count": 0,
            "cid": "1117",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
        },
        {
            "name": "Other",
            "residue": "other",
            "count": 0
        }
    ],
    "composition_expanded": [
        {
            "name": "Alditol",
            "residue": "aldi",
            "count": 0
        },
        {
            "name": "6-Deoxy-Hexose",
            "residue": "dhex",
            "count": 2,
            "cid": "840",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
        },
        {
            "name": "6-Deoxy-Hexitol",
            "residue": "dhex+aldi",
            "count": 0
        },
        {
            "name": "L-Fucose",
            "residue": "fuc",
            "count": 2,
            "cid": "17106",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/17106"
        },
        {
            "name": "L-Fucitol",
            "residue": "fuc+aldi",
            "count": 0,
            "cid": "445724",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/445724"
        },
        {
            "name": "D-Galactose",
            "residue": "gal",
            "count": 1,
            "cid": "6036",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6036"
        },
        {
            "name": "D-Galacitol",
            "residue": "gal+aldi",
            "count": 0,
            "cid": "11850",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/11850"
        },
        {
            "name": "D-Galactosamine",
            "residue": "galn",
            "count": 0,
            "cid": "24154",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/24154"
        },
        {
            "name": "N-Acetyl-D-galactosamine",
            "residue": "galnac",
            "count": 0,
            "cid": "35717",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/35717"
        },
        {
            "name": "N-Acetylgalactosaminitol",
            "residue": "galnac+aldi",
            "count": 0,
            "cid": "165880",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/165880"
        },
        {
            "name": "D-Glucose",
            "residue": "glc",
            "count": 0,
            "cid": "5793",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/5793"
        },
        {
            "name": "D-Glucitol",
            "residue": "glc+aldi",
            "count": 0,
            "cid": "5780",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/5780"
        },
        {
            "name": "D-Glucuronic acid",
            "residue": "glca",
            "count": 0,
            "cid": "94715",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/94715"
        },
        {
            "name": "D-Glucosamine",
            "residue": "glcn",
            "count": 0,
            "cid": "439213",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439213"
        },
        {
            "name": "N-Acetyl-D-Glucosamine",
            "residue": "glcnac",
            "count": 1,
            "cid": "439174",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439174"
        },
        {
            "name": "N-Acetyl-D-glucosaminitol",
            "residue": "glcnac+aldi",
            "count": 0,
            "cid": "165206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/165206"
        },
        {
            "name": "Hexose",
            "residue": "hex",
            "count": 1,
            "cid": "206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206"
        },
        {
            "name": "N-Acetylhexosamine",
            "residue": "hexnac",
            "count": 1,
            "cid": "899",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
        },
        {
            "name": "Hexuronic Acid",
            "residue": "hexa",
            "count": 0,
            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
        },
        {
            "name": "2-Amino-2-Deoxy-Hexose",
            "residue": "hexn",
            "count": 0,
            "cid": "739",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739"
        },
        {
            "name": "Hexitol",
            "residue": "hex+aldi",
            "count": 0,
            "cid": "453",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/453"
        },
        {
            "name": "N-Acetylhexosaminitol",
            "residue": "hexnac+aldi",
            "count": 0
        },
        {
            "name": "L-Iduronic acid",
            "residue": "idoa",
            "count": 0,
            "cid": "441039",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/441039"
        },
        {
            "name": "D-Mannose",
            "residue": "man",
            "count": 0,
            "cid": "18950",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/18950"
        },
        {
            "name": "D-Mannitol",
            "residue": "man+aldi",
            "count": 0,
            "cid": "6251",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6251"
        },
        {
            "name": "D-Mannosamine",
            "residue": "mann",
            "count": 0,
            "cid": "440049",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440049"
        },
        {
            "name": "N-Acetyl-D-mannosamine",
            "residue": "mannac",
            "count": 0,
            "cid": "439281",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439281"
        },
        {
            "name": "Methyl",
            "residue": "me",
            "count": 0
        },
        {
            "name": "N-Acetyl-Neuraminic Acid",
            "residue": "neuac",
            "count": 0,
            "cid": "439197",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
        },
        {
            "name": "N-Glycolyl-Neuraminic Acid",
            "residue": "neugc",
            "count": 0,
            "cid": "440001",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
        },
        {
            "name": "Phosphate",
            "residue": "p",
            "count": 0,
            "cid": "1061",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
        },
        {
            "name": "Pentose",
            "residue": "pent",
            "count": 0,
            "cid": "229",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
        },
        {
            "name": "Sulfate",
            "residue": "s",
            "count": 0,
            "cid": "1117",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
        },
        {
            "name": "Sialic acid",
            "residue": "sia",
            "count": 0,
            "cid": "906",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/906"
        },
        {
            "name": "Other",
            "residue": "other",
            "count": 0
        },
        {
            "name": "Floating substituent",
            "residue": "x",
            "count": 0
        },
        {
            "name": "D-Xylose",
            "residue": "xyl",
            "count": 0,
            "cid": "135191",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/135191"
        },
        {
            "name": "Ketodeoxynononic acid",
            "residue": "kdn",
            "count": 0,
            "cid": "13991616",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/13991616"
        }
    ],
    "motifs": [],
    "interactions": [],
    "expression": [],
    "subsumption": [
        {
            "related_accession": "G17991UN",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G18219LU",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G22979OY",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G23571FL",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G36592JW",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G57824FG",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G68835PZ",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G81216LK",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G82458HY",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G82458HY",
            "relationship": "basecomposition"
        },
        {
            "related_accession": "G22979OY",
            "relationship": "composition"
        },
        {
            "related_accession": "G00052MO",
            "relationship": "descendant"
        },
        {
            "related_accession": "G02485MC",
            "relationship": "descendant"
        },
        {
            "related_accession": "G49963IQ",
            "relationship": "descendant"
        },
        {
            "related_accession": "G82236PZ",
            "relationship": "descendant"
        },
        {
            "related_accession": "G96179XD",
            "relationship": "descendant"
        },
        {
            "related_accession": "G00052MO",
            "relationship": "fullydetermined"
        },
        {
            "related_accession": "G82236PZ",
            "relationship": "fullydetermined"
        },
        {
            "related_accession": "G00052MO",
            "relationship": "leaf"
        },
        {
            "related_accession": "G82236PZ",
            "relationship": "leaf"
        },
        {
            "related_accession": "G23571FL",
            "relationship": "subsumedby"
        },
        {
            "related_accession": "G68835PZ",
            "relationship": "subsumedby"
        },
        {
            "related_accession": "G02485MC",
            "relationship": "subsumes"
        },
        {
            "related_accession": "G81216LK",
            "relationship": "topology"
        }
    ],
    "names": [],
    "keywords": [],
    "glycotree_pathways": [],
    "dictionary": {}
}