{
    "glytoucan_ac": "G00061MO",
    "number_monosaccharides": 4,
    "number_monosaccharides_suffix": "",
    "mass": 707.25,
    "mass_pme": 903.47,
    "tool_support": {
        "gnome_glygen_oglycans": "no",
        "sandbox": "no",
        "gnome_glygen": "yes",
        "gnome": "yes",
        "gnome_glycotree_oglycans": "no",
        "pdb": "yes",
        "gnome_glygen_nglycans": "no",
        "gnome_glycotree_nglycans": "no"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "beta-D-GalpNAc-(1->3)-alpha-D-Galp-(1->4)-beta-D-Galp-(1->4)-beta-D-Glcp-(1->",
    "wurcs": "WURCS=2.0/4,4,3/[a2122h-1b_1-5][a2112h-1b_1-5][a2112h-1a_1-5][a2112h-1b_1-5_2*NCC/3=O]/1-2-3-4/a4-b1_b4-c1_c3-d1",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2b:b-dgal-HEX-1:5 3b:a-dgal-HEX-1:5 4b:b-dgal-HEX-1:5 5s:n-acetyl LIN 1:1o(4+1)2d 2:2o(4+1)3d 3:3o(3+1)4d 4:4d(2+1)5n",
    "inchi": "InChI=1S/C26H45NO21/c1-6(32)27-11-14(35)12(33)7(2-28)43-24(11)48-22-13(34)8(3-29)44-26(19(22)40)47-21-10(5-31)45-25(18(39)16(21)37)46-20-9(4-30)42-23(41)17(38)15(20)36/h7-26,28-31,33-41H,2-5H2,1H3,(H,27,32)/t7-,8-,9-,10-,11-,12+,13+,14-,15-,16-,17-,18-,19-,20-,21+,22+,23-,24+,25+,26-/m1/s1",
    "glycam": "DGalpNAcb1-3DGalpa1-4DGalpb1-4DGlcpb1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H]([C@H]([C@H](O[C@H]1O[C@H]2[C@H]([C@H](O[C@@H]([C@@H]2O)O[C@H]3[C@H](O[C@H]([C@@H]([C@H]3O)O)O[C@@H]4[C@H](O[C@H]([C@@H]([C@H]4O)O)O)CO)CO)CO)O)CO)O)O",
    "inchi_key": {
        "key": "JMPCSGHAAIDYIN-NYIFDZJOSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/JMPCSGHAAIDYIN-NYIFDZJOSA-N"
    },
    "byonic": "Hex(3)HexNAc(1) % 707.248407805",
    "gwb": "freeEnd--1b1D-Glc,p--4b1D-Gal,p--4a1D-Gal,p--3b1D-GalNAc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "Direct",
            "evidence": [
                {
                    "database": "GlyCosmos",
                    "id": "G00061MO",
                    "url": "https://glycosmos.org/glycans/show/G00061MO"
                },
                {
                    "database": "GlyTouCan",
                    "id": "G00061MO",
                    "url": "https://glytoucan.org/Structures/Glycans/G00061MO"
                }
            ],
            "common_name": "Human"
        },
        {
            "taxid": 10090,
            "name": "Mus musculus",
            "annotation_category": "Direct",
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                {
                    "database": "GlyCosmos",
                    "id": "G00061MO",
                    "url": "https://glycosmos.org/glycans/show/G00061MO"
                }
            ],
            "common_name": "Mouse"
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        {
            "taxid": 10114,
            "name": "Rattus",
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                    "database": "GlyCosmos",
                    "id": "G00061MO",
                    "url": "https://glycosmos.org/glycans/show/G00061MO"
                }
            ],
            "common_name": "rats"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "Glycosphingolipid"
            },
            "subtype": {
                "name": "Globo series"
            }
        }
    ],
    "enzyme": [],
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            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
        },
        {
            "name": "Hexuronic Acid",
            "residue": "hexa",
            "count": 0,
            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
        },
        {
            "name": "2-Amino-2-Deoxy-Hexose",
            "residue": "hexn",
            "count": 0,
            "cid": "739",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739"
        },
        {
            "name": "Hexitol",
            "residue": "hex+aldi",
            "count": 0,
            "cid": "453",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/453"
        },
        {
            "name": "N-Acetylhexosaminitol",
            "residue": "hexnac+aldi",
            "count": 0
        },
        {
            "name": "L-Iduronic acid",
            "residue": "idoa",
            "count": 0,
            "cid": "441039",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/441039"
        },
        {
            "name": "D-Mannose",
            "residue": "man",
            "count": 0,
            "cid": "18950",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/18950"
        },
        {
            "name": "D-Mannitol",
            "residue": "man+aldi",
            "count": 0,
            "cid": "6251",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6251"
        },
        {
            "name": "D-Mannosamine",
            "residue": "mann",
            "count": 0,
            "cid": "440049",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440049"
        },
        {
            "name": "N-Acetyl-D-mannosamine",
            "residue": "mannac",
            "count": 0,
            "cid": "439281",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439281"
        },
        {
            "name": "Methyl",
            "residue": "me",
            "count": 0
        },
        {
            "name": "N-Acetyl-Neuraminic Acid",
            "residue": "neuac",
            "count": 0,
            "cid": "439197",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
        },
        {
            "name": "N-Glycolyl-Neuraminic Acid",
            "residue": "neugc",
            "count": 0,
            "cid": "440001",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
        },
        {
            "name": "Phosphate",
            "residue": "p",
            "count": 0,
            "cid": "1061",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
        },
        {
            "name": "Pentose",
            "residue": "pent",
            "count": 0,
            "cid": "229",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
        },
        {
            "name": "Sulfate",
            "residue": "s",
            "count": 0,
            "cid": "1117",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
        },
        {
            "name": "Sialic acid",
            "residue": "sia",
            "count": 0,
            "cid": "906",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/906"
        },
        {
            "name": "Other",
            "residue": "other",
            "count": 0
        },
        {
            "name": "Floating substituent",
            "residue": "x",
            "count": 0
        },
        {
            "name": "D-Xylose",
            "residue": "xyl",
            "count": 0,
            "cid": "135191",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/135191"
        },
        {
            "name": "Ketodeoxynononic acid",
            "residue": "kdn",
            "count": 0,
            "cid": "13991616",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/13991616"
        }
    ],
    "motifs": [
        {
            "id": "GGM.000070",
            "name": "Globoside",
            "synonym": "Cytolipin K; Gb4; Globoside I; Globotetraosylceramide; P antigen",
            "keywords": "Glycolipid"
        }
    ],
    "interactions": [],
    "expression": [],
    "subsumption": [
        {
            "related_accession": "G08590QR",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G13769ML",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G15216HY",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G24499XE",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G29964CE",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G33233SN",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G48148TW",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G49958GL",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G52725AC",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G53509NM",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G81647RY",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G82558IX",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G82907DV",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G82930TB",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G86170TS",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G99312NX",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G08590QR",
            "relationship": "basecomposition"
        },
        {
            "related_accession": "G53509NM",
            "relationship": "composition"
        },
        {
            "related_accession": "G24499XE",
            "relationship": "subsumedby"
        },
        {
            "related_accession": "G81647RY",
            "relationship": "topology"
        }
    ],
    "names": [
        {
            "name": "Globoside",
            "domain": "Semantic Name"
        }
    ],
    "keywords": [
        "fully_determined"
    ],
    "glycotree_pathways": [],
    "dictionary": {
        "term": "Globotetraosylceramide",
        "term_in_sentence": "Here we report that PUGNAc treatment provokes globotetraosylceramide (Gb4Cer) accumulation in human umbilical vein endothelial cells (HUVEC).[PMID:28392398]",
        "publication": "28392398|25787850|23471986|2559078|32455599|2663859|32796033|3517161|20082214|30853699|15279935|26104834|20732996|30224239|31142708|30338271|25033755|30413389|7104376|29866658|24983355|6794884|31361021|21740983|33168837|31752441|28123520|29068380|24841197|27932383|28535204|31691795|27916888|27558838|10227680|26464281|22348006|22279060|25156739|32314902|22718629|23701631|23906628|8613382|23555772|20059899|7868240",
        "definition": "An amino tetrasaccharide consisting of 2-acetamido-2-deoxy-\u03b2-D-galactopyranosyl, \u03b1-D-galactopyranosyl, \u03b2-D-galactopyranosyl, and \u03b2-D-glucopyranose units joined together in sequence by (1\u21923), (1\u21924), and (1\u21924) glycosidic linkages, respectively.[CHEBI:145578]",
        "term_xref": "GlycoMotif:GGM.000070|GlycoEpitope:EP0073|CID:5288889|CHEBI:145578|SugarBind_Ligand:18|MeSH:C024032",
        "synonymns": [
            "Gb4Cer",
            "Gb4",
            "Globoside",
            "P antigen",
            "Cytolipin K"
        ],
        "function": "",
        "disease_associations": "Actinomycosis[SugarBind_Ligand:18]|Pyelonephritis[SugarBind_Ligand:18]|Acute cystitis[SugarBind_Ligand:18]|Shigellosis[SugarBind_Ligand:18]|Lazy leukocyte syndrome[SugarBind_Ligand:18]|Influenza[SugarBind_Ligand:18]",
        "wikipedia": "",
        "essentials_of_glycobiology": "Chapter 11",
        "evidence": [
            {
                "id": "GSD000088",
                "url": "https://wiki.glygen.org/Glycan_structure_dictionary#GSD000088",
                "database": "Glycan Structure Dictionary"
            }
        ]
    }
}