{
    "glytoucan_ac": "G05191RT",
    "number_monosaccharides": 12,
    "number_monosaccharides_suffix": "",
    "mass": 2282.84,
    "mass_pme": 2843.47,
    "tool_support": {
        "sandbox": "no",
        "pdb": "no",
        "gnome": "yes",
        "gnome_glygen": "yes",
        "gnome_glygen_nglycans": "no",
        "gnome_glygen_oglycans": "no",
        "gnome_glycotree_nglycans": "no",
        "gnome_glycotree_oglycans": "no"
    },
    "fully_determined": "",
    "missing_score": 902,
    "glycan_type": "Saccharide",
    "iupac": "",
    "iupac_compact": "NeuAc?2-4Gal?1-4GlcNAc?1-2Man?1-3(Gal?1-4GlcNAc?1-2(GlcNAc?1-6)Man?1-6)Man?1-4GlcNAc?1-4(Fuc?1-6)GlcNAc+aldi",
    "iupac_condensed": "",
    "wurcs": "WURCS=2.0/6,12,11/[h2122h_2*NCC/3=O][a2122h-1x_1-5_2*NCC/3=O][a1122h-1x_1-5][a2112h-1x_1-5][Aad21122h-2x_2-6_5*NCC/3=O][a1221m-1x_1-5]/1-2-3-3-2-4-5-3-2-4-2-6/a4-b1_a6-l1_b4-c1_c3-d1_c6-h1_d2-e1_e4-f1_f4-g2_h2-i1_h6-k1_i4-j1",
    "glycoct": "RES 1b:o-dglc-HEX-0:0|1:aldi 2s:n-acetyl 3b:x-dglc-HEX-1:5 4s:n-acetyl 5b:x-dman-HEX-1:5 6b:x-dman-HEX-1:5 7b:x-dglc-HEX-1:5 8s:n-acetyl 9b:x-dgal-HEX-1:5 10b:x-dgro-dgal-NON-2:6|1:a|2:keto|3:d 11s:n-acetyl 12b:x-dman-HEX-1:5 13b:x-dglc-HEX-1:5 14s:n-acetyl 15b:x-dgal-HEX-1:5 16b:x-dglc-HEX-1:5 17s:n-acetyl 18b:x-lgal-HEX-1:5|6:d LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:9o(4+2)10d 10:10d(5+1)11n 11:5o(6+1)12d 12:12o(2+1)13d 13:13d(2+1)14n 14:13o(4+1)15d 15:12o(6+1)16d 16:16d(2+1)17n 17:1o(6+1)18d",
    "inchi": "InChI=1S/C87H146N6O63/c1-21-46(112)58(124)63(129)80(138-21)135-18-31(111)67(47(113)28(9-94)88-22(2)103)148-77-43(91-25(5)106)55(121)70(37(16-101)142-77)151-83-66(132)73(53(119)40(146-83)20-137-84-74(61(127)52(118)39(147-84)19-136-76-42(90-24(4)105)54(120)49(115)32(11-96)139-76)153-78-44(92-26(6)107)56(122)68(35(14-99)143-78)149-81-64(130)59(125)50(116)33(12-97)140-81)152-85-75(60(126)51(117)34(13-98)141-85)154-79-45(93-27(7)108)57(123)69(36(15-100)144-79)150-82-65(131)62(128)71(38(17-102)145-82)155-87(86(133)134)8-29(109)41(89-23(3)104)72(156-87)48(114)30(110)10-95/h21,28-85,94-102,109-132H,8-20H2,1-7H3,(H,88,103)(H,89,104)(H,90,105)(H,91,106)(H,92,107)(H,93,108)(H,133,134)/t21-,28-,29-,30+,31+,32+,33+,34+,35+,36+,37+,38+,39+,40+,41+,42+,43+,44+,45+,46+,47+,48+,49+,50-,51+,52+,53+,54+,55+,56+,57+,58+,59-,60-,61-,62+,63-,64+,65+,66-,67+,68+,69+,70+,71-,72+,73-,74-,75-,76?,77?,78?,79?,80?,81?,82?,83?,84?,85?,87?/m0/s1",
    "glycam": "",
    "smiles_isomeric": "C[C@H]1[C@H]([C@H]([C@@H](C(O1)OC[C@H]([C@H]([C@@H]([C@H](CO)NC(=O)C)O)OC2[C@@H]([C@H]([C@@H]([C@H](O2)CO)OC3[C@H]([C@H]([C@@H]([C@H](O3)COC4[C@H]([C@H]([C@@H]([C@H](O4)COC5[C@@H]([C@H]([C@@H]([C@H](O5)CO)O)O)NC(=O)C)O)O)OC6[C@@H]([C@H]([C@@H]([C@H](O6)CO)OC7[C@@H]([C@H]([C@H]([C@H](O7)CO)O)O)O)O)NC(=O)C)O)OC8[C@H]([C@H]([C@@H]([C@H](O8)CO)O)O)OC9[C@@H]([C@H]([C@@H]([C@H](O9)CO)OC1[C@@H]([C@H]([C@H]([C@H](O1)CO)OC1(C[C@@H]([C@H]([C@@H](O1)[C@@H]([C@@H](CO)O)O)NC(=O)C)O)C(=O)O)O)O)O)NC(=O)C)O)O)NC(=O)C)O)O)O)O",
    "inchi_key": {
        "key": "XAGPSHCPPUUWIN-XVYHHNNMSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/XAGPSHCPPUUWIN-XVYHHNNMSA-N"
    },
    "byonic": "",
    "gwb": "redEnd--?D-GlcNAc,o(--4?1D-GlcNAc,p--4?1D-Man,p(--3?1D-Man,p--2?1D-GlcNAc,p--4?1D-Gal,p--4?2D-NeuAc,p)--6?1D-Man,p(--2?1D-GlcNAc,p--4?1D-Gal,p)--6?1D-GlcNAc,p)--6?1L-Fuc,p$MONO,Und,0,0,redEnd",
    "species": [
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "PubMed",
                    "id": "39988192",
                    "url": "https://glygen.org/publication/PubMed/39988192"
                },
                {
                    "database": "GlyGen",
                    "id": "GLY_001408",
                    "url": "https://data.glygen.org/GLY_001408"
                }
            ],
            "common_name": "Human",
            "glygen_name": "Human",
            "reference_species": "Homo sapiens [9606]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Alditol-reduced"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Complex",
                "url": ""
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Core-fucosylated"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Triantennary"
            }
        }
    ],
    "enzyme": [],
    "crossref": [
        {
            "id": "176511660",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/176511660",
            "database": "PubChem Compound",
            "categories": [
                "Chemical"
            ]
        },
        {
            "id": "518302073",
            "url": "https://pubchem.ncbi.nlm.nih.gov/substance/518302073",
            "database": "PubChem Substance",
            "categories": [
                "Chemical"
            ]
        },
        {
            "id": "G05191RT",
            "url": "https://glytoucan.org/Structures/Glycans/G05191RT",
            "database": "GlyTouCan",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "G05191RT",
            "url": "https://gnome.glyomics.org/restrictions/GlyGen.StructureBrowser.html?focus=G05191RT",
            "database": "GNOme",
            "categories": [
                "Glycomics",
                "Ontology and Dictionary"
            ]
        },
        {
            "id": "G05191RT",
            "url": "https://glycosmos.org/glycans/show/G05191RT",
            "database": "GlyCosmos",
            "categories": [
                "Glycoproteomics",
                "Glycomics"
            ]
        }
    ],
    "glycoprotein": [],
    "publication": [
        {
            "title": "Integrated Multiomics Reveals Alterations in Paucimannose and Complex Type N-Glycans in Cardiac Tissue of Patients with COVID-19.",
            "journal": "Molecular & cellular proteomics : MCP",
            "date": "2025-04",
            "authors": "Subramanian Sabarinath Peruvemba, Wojtkiewicz Melinda, Yu Fang, Castro Chase, Schuette Erin N, Rodriguez-Paar Jocelyn, Churko Jared, Renavikar Pranav, Anderson Daniel, Mahr Claudius, Gundry Rebekah L",
            "evidence": [
                {
                    "database": "TableMaker",
                    "id": "TG1131507",
                    "url": "https://glygen.ccrc.uga.edu/tablemaker/data/dataset/TG1131507"
                },
                {
                    "database": "GlyTouCan",
                    "id": "G05191RT",
                    "url": "https://glytoucan.org/Structures/Glycans/G05191RT"
                }
            ],
            "reference": [
                {
                    "type": "PubMed",
                    "id": "39988192",
                    "url": "https://glygen.org/publication/PubMed/39988192"
                }
            ]
        }
    ],
    "residues": [],
    "composition": [
        {
            "name": "6-Deoxy-Hexose",
            "residue": "dhex",
            "count": 1,
            "cid": "840",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
        },
        {
            "name": "Hexose",
            "residue": "hex",
            "count": 5,
            "cid": "206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206"
        },
        {
            "name": "N-Acetylhexosamine",
            "residue": "hexnac",
            "count": 5,
            "cid": "899",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
        },
        {
            "name": "Hexuronic Acid",
            "residue": "hexa",
            "count": 0,
            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
        },
        {
            "name": "2-Amino-2-Deoxy-Hexose",
            "residue": "hexn",
            "count": 0,
            "cid": "739",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739"
        },
        {
            "name": "N-Acetyl-Neuraminic Acid",
            "residue": "neuac",
            "count": 1,
            "cid": "439197",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
        },
        {
            "name": "N-Glycolyl-Neuraminic Acid",
            "residue": "neugc",
            "count": 0,
            "cid": "440001",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
        },
        {
            "name": "Phosphate",
            "residue": "p",
            "count": 0,
            "cid": "1061",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
        },
        {
            "name": "Pentose",
            "residue": "pent",
            "count": 0,
            "cid": "229",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
        },
        {
            "name": "Sulfate",
            "residue": "s",
            "count": 0,
            "cid": "1117",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
        },
        {
            "name": "Other",
            "residue": "other",
            "count": 0
        }
    ],
    "composition_expanded": [
        {
            "name": "Alditol",
            "residue": "aldi",
            "count": 1
        },
        {
            "name": "6-Deoxy-Hexose",
            "residue": "dhex",
            "count": 1,
            "cid": "840",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
        },
        {
            "name": "6-Deoxy-Hexitol",
            "residue": "dhex+aldi",
            "count": 0
        },
        {
            "name": "L-Fucose",
            "residue": "fuc",
            "count": 1,
            "cid": "17106",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/17106"
        },
        {
            "name": "L-Fucitol",
            "residue": "fuc+aldi",
            "count": 0,
            "cid": "445724",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/445724"
        },
        {
            "name": "D-Galactose",
            "residue": "gal",
            "count": 2,
            "cid": "6036",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6036"
        },
        {
            "name": "D-Galactitol",
            "residue": "gal+aldi",
            "count": 0,
            "cid": "11850",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/11850"
        },
        {
            "name": "N-Acetyl-D-galactosamine",
            "residue": "galnac",
            "count": 0,
            "cid": "35717",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/35717"
        },
        {
            "name": "N-Acetylgalactosaminitol",
            "residue": "galnac+aldi",
            "count": 0,
            "cid": "165880",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/165880"
        },
        {
            "name": "D-Galacturonic Acid",
            "residue": "gala",
            "count": 0,
            "cid": "439215",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439215"
        },
        {
            "name": "D-Glucose",
            "residue": "glc",
            "count": 0,
            "cid": "5793",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/5793"
        },
        {
            "name": "D-Glucitol",
            "residue": "glc+aldi",
            "count": 0,
            "cid": "5780",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/5780"
        },
        {
            "name": "D-Glucuronic acid",
            "residue": "glca",
            "count": 0,
            "cid": "94715",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/94715"
        },
        {
            "name": "D-Glucosamine",
            "residue": "glcn",
            "count": 0,
            "cid": "439213",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439213"
        },
        {
            "name": "N-Acetyl-D-Glucosamine",
            "residue": "glcnac",
            "count": 5,
            "cid": "439174",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439174"
        },
        {
            "name": "N-Acetyl-D-glucosaminitol",
            "residue": "glcnac+aldi",
            "count": 1,
            "cid": "165206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/165206"
        },
        {
            "name": "Hexose",
            "residue": "hex",
            "count": 5,
            "cid": "206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206"
        },
        {
            "name": "N-Acetylhexosamine",
            "residue": "hexnac",
            "count": 5,
            "cid": "899",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
        },
        {
            "name": "Hexuronic Acid",
            "residue": "hexa",
            "count": 0,
            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
        },
        {
            "name": "2-Amino-2-Deoxy-Hexose",
            "residue": "hexn",
            "count": 0,
            "cid": "739",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739"
        },
        {
            "name": "Hexitol",
            "residue": "hex+aldi",
            "count": 0,
            "cid": "453",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/453"
        },
        {
            "name": "N-Acetylhexosaminitol",
            "residue": "hexnac+aldi",
            "count": 1
        },
        {
            "name": "L-Iduronic acid",
            "residue": "idoa",
            "count": 0,
            "cid": "441039",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/441039"
        },
        {
            "name": "D-Mannose",
            "residue": "man",
            "count": 3,
            "cid": "18950",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/18950"
        },
        {
            "name": "D-Mannitol",
            "residue": "man+aldi",
            "count": 0,
            "cid": "6251",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/6251"
        },
        {
            "name": "N-Acetyl-D-mannosamine",
            "residue": "mannac",
            "count": 0,
            "cid": "439281",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439281"
        },
        {
            "name": "Methyl",
            "residue": "me",
            "count": 0
        },
        {
            "name": "N-Acetyl-Neuraminic Acid",
            "residue": "neuac",
            "count": 1,
            "cid": "439197",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197"
        },
        {
            "name": "N-Glycolyl-Neuraminic Acid",
            "residue": "neugc",
            "count": 0,
            "cid": "440001",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001"
        },
        {
            "name": "Phosphate",
            "residue": "p",
            "count": 0,
            "cid": "1061",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061"
        },
        {
            "name": "Pentose",
            "residue": "pent",
            "count": 0,
            "cid": "229",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/229"
        },
        {
            "name": "Sulfate",
            "residue": "s",
            "count": 0,
            "cid": "1117",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117"
        },
        {
            "name": "Sialic acid",
            "residue": "sia",
            "count": 1
        },
        {
            "name": "Other",
            "residue": "other",
            "count": 0
        },
        {
            "name": "Floating substituent",
            "residue": "x",
            "count": 0
        },
        {
            "name": "D-Xylose",
            "residue": "xyl",
            "count": 0,
            "cid": "135191",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/135191"
        },
        {
            "name": "Ketodeoxynononic acid",
            "residue": "kdn",
            "count": 0,
            "cid": "13991616",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/13991616"
        }
    ],
    "motifs": [
        {
            "id": "GGM.000001",
            "name": "Type 2 LN",
            "synonym": "N-acetyllactosamine",
            "keywords": "LacNAc; N-acetyllactosamine; N-acetyllactosamine type 2"
        }
    ],
    "interactions": [],
    "expression": [
        {
            "category": "tissue",
            "tissue": {
                "name": "heart",
                "namespace": "UBERON",
                "id": "0000948",
                "url": "http://purl.obolibrary.org/obo/UBERON_0000948"
            },
            "cell_line": {},
            "abundance": "",
            "evidence": [
                {
                    "id": "TG1131507",
                    "database": "TableMaker",
                    "url": "https://glygen.ccrc.uga.edu/tablemaker/data/dataset/TG1131507"
                },
                {
                    "id": "39988192",
                    "database": "PubMed",
                    "url": "https://glygen.org/publication/PubMed/39988192"
                }
            ]
        }
    ],
    "tissue": [
        {
            "tissue": {
                "name": "heart",
                "namespace": "UBERON",
                "id": "0000948",
                "url": "http://purl.obolibrary.org/obo/UBERON_0000948"
            },
            "species": {
                "name": "Homo sapiens",
                "taxid": 9606,
                "reference_species": "Homo sapiens [9606]",
                "common_name": "Human",
                "glygen_name": "Human"
            },
            "evidence": [
                {
                    "id": "TG1131507",
                    "database": "TableMaker",
                    "url": "https://glygen.ccrc.uga.edu/tablemaker/data/dataset/TG1131507"
                },
                {
                    "id": "39988192",
                    "database": "PubMed",
                    "url": "https://glygen.org/publication/PubMed/39988192"
                }
            ]
        }
    ],
    "subsumption": [
        {
            "related_accession": "G05010JE",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G18842WA",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G76915EY",
            "relationship": "ancestor"
        },
        {
            "related_accession": "G18842WA",
            "relationship": "composition"
        },
        {
            "related_accession": "G05010JE",
            "relationship": "subsumedby"
        }
    ],
    "names": [],
    "keywords": [],
    "glycotree_pathways": [],
    "dictionary": {},
    "biomarkers": [],
    "section_stats": [
        {
            "table_id": "glycoprotein",
            "table_stats": [
                {
                    "field": "total",
                    "count": 0
                },
                {
                    "field": "total_sites",
                    "count": 0
                }
            ],
            "sort_fields": []
        },
        {
            "table_id": "expression",
            "table_stats": [
                {
                    "field": "total",
                    "count": 1
                },
                {
                    "field": "total_sites",
                    "count": 0
                }
            ],
            "sort_fields": [
                "category",
                "tissue.name",
                "tissue.namespace",
                "tissue.id",
                "tissue.url",
                "abundance"
            ]
        },
        {
            "table_id": "expression_tissue",
            "table_stats": [
                {
                    "field": "total",
                    "count": 1
                },
                {
                    "field": "total_sites",
                    "count": 0
                }
            ],
            "sort_fields": [
                "category",
                "tissue.name",
                "tissue.namespace",
                "tissue.id",
                "tissue.url",
                "abundance"
            ]
        },
        {
            "table_id": "expression_cell_line",
            "table_stats": [
                {
                    "field": "total",
                    "count": 0
                },
                {
                    "field": "total_sites",
                    "count": 0
                }
            ],
            "sort_fields": [
                "category",
                "abundance"
            ]
        },
        {
            "table_id": "publication",
            "table_stats": [
                {
                    "field": "total",
                    "count": 1
                }
            ],
            "sort_fields": [
                "title",
                "journal",
                "date",
                "authors"
            ]
        }
    ]
}