{
    "glytoucan_ac": "G04791QM",
    "number_monosaccharides": 9,
    "number_monosaccharides_suffix": "",
    "mass": 1769.63,
    "mass_pme": 2204.12,
    "tool_support": {
        "sandbox": "yes",
        "pdb": "yes",
        "gnome": "yes",
        "gnome_glygen": "yes",
        "gnome_glygen_nglycans": "yes",
        "gnome_glygen_oglycans": "no",
        "gnome_glycotree_nglycans": "yes",
        "gnome_glycotree_oglycans": "no"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "alpha-D-Neup5Ac-(2->6)-beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->2)-alpha-D-Manp-(1->3)[beta-D-GlcpNAc-(1->2)-alpha-D-Manp-(1->6)]-beta-D-Manp-(1->4)-beta-D-GlcpNAc-(1->4)-beta-D-GlcpNAc-(1->",
    "iupac_compact": "NeuAca2-6Galb1-4GlcNAcb1-2Mana1-3(GlcNAcb1-2Mana1-6)Manb1-4GlcNAcb1-4GlcNAcb",
    "iupac_condensed": "Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-",
    "wurcs": "WURCS=2.0/5,9,8/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O]/1-1-2-3-1-4-5-3-1/a4-b1_b4-c1_c3-d1_c6-h1_d2-e1_e4-f1_f6-g2_h2-i1",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:b-dglc-HEX-1:5 8s:n-acetyl 9b:b-dgal-HEX-1:5 10b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 11s:n-acetyl 12b:a-dman-HEX-1:5 13b:b-dglc-HEX-1:5 14s:n-acetyl LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(2+1)7d 7:7d(2+1)8n 8:7o(4+1)9d 9:9o(6+2)10d 10:10d(5+1)11n 11:5o(6+1)12d 12:12o(2+1)13d 13:13d(2+1)14n",
    "inchi": "InChI=1S/C67H111N5O49/c1-16(80)68-31-21(85)6-67(66(103)104,121-54(31)36(87)22(86)7-73)106-15-30-40(91)46(97)49(100)62(114-30)116-52-27(12-78)112-61(35(45(52)96)72-20(5)84)120-57-48(99)39(90)25(10-76)110-65(57)118-55-41(92)29(14-105-64-56(47(98)38(89)24(9-75)109-64)119-59-33(70-18(3)82)42(93)37(88)23(8-74)108-59)113-63(50(55)101)117-53-28(13-79)111-60(34(44(53)95)71-19(4)83)115-51-26(11-77)107-58(102)32(43(51)94)69-17(2)81/h21-65,73-79,85-102H,6-15H2,1-5H3,(H,68,80)(H,69,81)(H,70,82)(H,71,83)(H,72,84)(H,103,104)/t21-,22+,23+,24+,25+,26+,27+,28+,29+,30+,31+,32+,33+,34+,35+,36+,37+,38+,39+,40-,41+,42+,43+,44+,45+,46-,47-,48-,49+,50-,51+,52+,53+,54+,55-,56-,57-,58+,59-,60-,61-,62-,63-,64-,65+,67+/m0/s1",
    "glycam": "DNeup5Aca2-6DGalpb1-4DGlcpNAcb1-2DManpa1-3[DGlcpNAcb1-2DManpa1-6]DManpb1-4DGlcpNAcb1-4DGlcpNAcb1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H](C[C@@](O[C@H]1[C@@H]([C@@H](CO)O)O)(C(=O)O)OC[C@@H]2[C@@H]([C@@H]([C@H]([C@@H](O2)O[C@@H]3[C@H](O[C@H]([C@@H]([C@H]3O)NC(=O)C)O[C@H]4[C@H]([C@@H]([C@H](O[C@@H]4O[C@H]5[C@@H]([C@H](O[C@H]([C@H]5O)O[C@@H]6[C@H](O[C@H]([C@@H]([C@H]6O)NC(=O)C)O[C@@H]7[C@H](O[C@H]([C@@H]([C@H]7O)NC(=O)C)O)CO)CO)CO[C@@H]8[C@H]([C@H]([C@@H]([C@H](O8)CO)O)O)O[C@H]9[C@@H]([C@H]([C@@H]([C@H](O9)CO)O)O)NC(=O)C)O)CO)O)O)CO)O)O)O)O",
    "inchi_key": {
        "key": "IDNBCPDOXIQLSO-ZESRDZFBSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/IDNBCPDOXIQLSO-ZESRDZFBSA-N"
    },
    "byonic": "HexNAc(4)Hex(4)NeuAc(1) % 1769.63476575",
    "gwb": "freeEnd--1b1D-GlcNAc,p--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p--2b1D-GlcNAc,p--4b1D-Gal,p--6a2D-NeuAc,p)--6a1D-Man,p--2b1D-GlcNAc,p$MONO,Und,0,0,freeEnd",
    "species": [
        {
            "taxid": 9031,
            "name": "Gallus gallus",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "938",
                    "url": "https://glyconnect.expasy.org/browser/structures/938"
                },
                {
                    "database": "PubMed",
                    "id": "19270371",
                    "url": "https://glygen.org/publication/PubMed/19270371"
                },
                {
                    "database": "PubMed",
                    "id": "8370663",
                    "url": "https://glygen.org/publication/PubMed/8370663"
                }
            ],
            "common_name": "Chicken",
            "glygen_name": "Chicken",
            "reference_species": "Gallus gallus [9031]"
        },
        {
            "taxid": 9606,
            "name": "Homo sapiens",
            "annotation_category": "direct",
            "evidence": [
                {
                    "database": "GlyConnect",
                    "id": "938",
                    "url": "https://glyconnect.expasy.org/browser/structures/938"
                },
                {
                    "database": "GlyGen",
                    "id": "GLY_001412",
                    "url": "https://data.glygen.org/GLY_001412"
                },
                {
                    "database": "PubMed",
                    "id": "1694115",
                    "url": "https://glygen.org/publication/PubMed/1694115"
                },
                {
                    "database": "PubMed",
                    "id": "27617431",
                    "url": "https://glygen.org/publication/PubMed/27617431"
                },
                {
                    "database": "PubMed",
                    "id": "32841605",
                    "url": "https://glygen.org/publication/PubMed/32841605"
                },
                {
                    "database": "PubMed",
                    "id": "6192908",
                    "url": "https://glygen.org/publication/PubMed/6192908"
                },
                {
                    "database": "PubMed",
                    "id": "7511099",
                    "url": "https://glygen.org/publication/PubMed/7511099"
                },
                {
                    "database": "PubMed",
                    "id": "7679920",
                    "url": "https://glygen.org/publication/PubMed/7679920"
                },
                {
                    "database": "PubMed",
                    "id": "7686446",
                    "url": "https://glygen.org/publication/PubMed/7686446"
                },
                {
                    "database": "PubMed",
                    "id": "7785764",
                    "url": "https://glygen.org/publication/PubMed/7785764"
                },
                {
                    "database": "UniCarbKB",
                    "id": "GLY_000341",
                    "url": "http://data.glygen.org/GLY_000341"
                }
            ],
            "common_name": "Human",
            "glygen_name": "Human",
            "reference_species": "Homo sapiens [9606]"
        }
    ],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Biantennary"
            }
        },
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "Complex",
                "url": ""
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprotkb/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "Alg1",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P26572-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT1",
            "gene_link": "https://www.uniprot.org/uniprotkb/P26572",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P27808-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat1",
            "gene_link": "https://www.uniprot.org/uniprotkb/P27808",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60512-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4GALT3",
            "gene_link": "https://www.uniprot.org/uniprotkb/O60512",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "O60909-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4GALT2",
            "gene_link": "https://www.uniprot.org/uniprotkb/O60909",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15291-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4GALT1",
            "gene_link": "https://www.uniprot.org/uniprotkb/P15291",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "P15535-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Beta-1,4-galactosyltransferase 1",
            "gene": "B4galt1",
            "gene_link": "https://www.uniprot.org/uniprotkb/P15535",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q91YY2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Beta-1,4-galactosyltransferase 3",
            "gene": "B4galt3",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q91YY2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Beta-1,4-galactosyltransferase 2",
            "gene": "B4galt2",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9Z2Y2",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q10469-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "MGAT2",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q10469",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G04791QM",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G04791QM"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921V5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G04791QM",
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase",
            "gene": "Mgat2",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q921V5",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
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