{
    "glytoucan_ac": "G00855BY",
    "number_monosaccharides": 11,
    "number_monosaccharides_suffix": "",
    "mass": 1882.64,
    "mass_pme": 2373.19,
    "tool_support": {
        "sandbox": "yes",
        "pdb": "no",
        "gnome": "yes",
        "gnome_glygen": "yes",
        "gnome_glygen_nglycans": "no",
        "gnome_glygen_oglycans": "no",
        "gnome_glycotree_nglycans": "yes",
        "gnome_glycotree_oglycans": "no"
    },
    "fully_determined": "yes",
    "missing_score": 0,
    "glycan_type": "Saccharide",
    "iupac": "alpha-D-Manp-(1->4)-alpha-D-Manp-(1->4)-alpha-D-Manp-(1->3)[alpha-D-Manp-(1->4)-alpha-D-Manp-(1->3)[alpha-D-Manp-(1->4)-alpha-D-Manp-(1->6)]-alpha-D-Manp-(1->6)]-beta-D-Manp-(1->4)-beta-D-GlcpNAc-(1->4)-beta-D-GlcpNAc-(1->",
    "iupac_compact": "Mana1-4Mana1-4Mana1-3(Mana1-4Mana1-3(Mana1-4Mana1-6)Mana1-6)Manb1-4GlcNAcb1-4GlcNAcb",
    "iupac_condensed": "Man(a1-4)Man(a1-4)Man(a1-3)[Man(a1-4)Man(a1-3)[Man(a1-4)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-",
    "wurcs": "WURCS=2.0/3,11,10/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5]/1-1-2-3-3-3-3-3-3-3-3/a4-b1_b4-c1_c3-d1_c6-g1_d4-e1_e4-f1_g3-h1_g6-j1_h4-i1_j4-k1",
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:a-dman-HEX-1:5 8b:a-dman-HEX-1:5 9b:a-dman-HEX-1:5 10b:a-dman-HEX-1:5 11b:a-dman-HEX-1:5 12b:a-dman-HEX-1:5 13b:a-dman-HEX-1:5 LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(4+1)7d 7:7o(4+1)8d 8:5o(6+1)9d 9:9o(3+1)10d 10:10o(4+1)11d 11:9o(6+1)12d 12:12o(4+1)13d",
    "inchi": "InChI=1S/C70H118N2O56/c1-14(82)71-27-34(89)52(19(6-76)110-60(27)107)121-61-28(72-15(2)83)35(90)53(20(7-77)114-61)122-70-51(106)59(128-69-49(104)42(97)57(24(11-81)118-69)126-67-47(102)40(95)55(22(9-79)116-67)124-65-44(99)37(92)30(85)17(4-74)112-65)33(88)26(120-70)13-109-63-50(105)58(127-68-48(103)41(96)56(23(10-80)117-68)125-66-45(100)38(93)31(86)18(5-75)113-66)32(87)25(119-63)12-108-62-46(101)39(94)54(21(8-78)115-62)123-64-43(98)36(91)29(84)16(3-73)111-64/h16-70,73-81,84-107H,3-13H2,1-2H3,(H,71,82)(H,72,83)/t16-,17-,18-,19-,20-,21-,22-,23-,24-,25-,26-,27-,28-,29-,30-,31-,32-,33-,34-,35-,36+,37+,38+,39-,40-,41-,42-,43+,44+,45+,46+,47+,48+,49+,50+,51+,52-,53-,54-,55-,56-,57-,58+,59+,60-,61+,62+,63+,64-,65-,66-,67-,68-,69-,70+/m1/s1",
    "glycam": "DManpa1-4DManpa1-3[DManpa1-4DManpa1-6]DManpa1-6[DManpa1-4DManpa1-4DManpa1-3]DManpb1-4DGlcpNAcb1-4DGlcpNAcb1-OH",
    "smiles_isomeric": "CC(=O)N[C@@H]1[C@H]([C@@H]([C@H](O[C@H]1O)CO)O[C@H]2[C@@H]([C@H]([C@@H]([C@H](O2)CO)O[C@H]3[C@H]([C@H]([C@@H]([C@H](O3)CO[C@@H]4[C@H]([C@H]([C@@H]([C@H](O4)CO[C@@H]5[C@H]([C@H]([C@@H]([C@H](O5)CO)O[C@@H]6[C@H]([C@H]([C@@H]([C@H](O6)CO)O)O)O)O)O)O)O[C@@H]7[C@H]([C@H]([C@@H]([C@H](O7)CO)O[C@@H]8[C@H]([C@H]([C@@H]([C@H](O8)CO)O)O)O)O)O)O)O)O[C@@H]9[C@H]([C@H]([C@@H]([C@H](O9)CO)O[C@@H]1[C@H]([C@H]([C@@H]([C@H](O1)CO)O[C@@H]1[C@H]([C@H]([C@@H]([C@H](O1)CO)O)O)O)O)O)O)O)O)O)NC(=O)C)O",
    "inchi_key": {
        "key": "FFCVHCXQYMZNBV-FRGQJGDUSA-N",
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/FFCVHCXQYMZNBV-FRGQJGDUSA-N"
    },
    "byonic": "HexNAc(2)Hex(9) % 1882.64472141",
    "gwb": "freeEnd--1b1D-GlcNAc,p--4b1D-GlcNAc,p--4b1D-Man,p(--3a1D-Man,p--4a1D-Man,p--4a1D-Man,p)--6a1D-Man,p(--3a1D-Man,p--4a1D-Man,p)--6a1D-Man,p--4a1D-Man,p$MONO,Und,0,0,freeEnd",
    "species": [],
    "classification": [
        {
            "type": {
                "name": "N-linked",
                "url": ""
            },
            "subtype": {
                "name": "High mannose",
                "url": ""
            }
        }
    ],
    "enzyme": [
        {
            "uniprot_canonical_ac": "P42867-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "Dpagt1",
            "gene_link": "https://www.uniprot.org/uniprotkb/P42867",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H3H5-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase",
            "gene": "DPAGT1",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9H3H5",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8BML3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Glt28d2",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q8BML3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q96F25-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "ALG14",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q96F25",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D081-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14",
            "gene": "Alg14",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9D081",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9D8C3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "Alg13",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9D8C3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
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                    "id": "G00855BY",
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                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9NP73-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG13",
            "gene": "ALG13",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9NP73",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q921Q3-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
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            "gene_link": "https://www.uniprot.org/uniprotkb/Q921Q3",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9BT22-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase",
            "gene": "ALG1",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9BT22",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9DBE8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "Alg2",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9DBE8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q9H553-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2",
            "gene": "ALG2",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q9H553",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q8K2A8-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "Dol-P-Man:Man(5)GlcNAc(2)-PP-Dol alpha-1,3-mannosyltransferase",
            "gene": "Alg3",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q8K2A8",
            "tax_id": 10090,
            "tax_name": "Mus musculus",
            "tax_common_name": "Mouse",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        },
        {
            "uniprot_canonical_ac": "Q92685-1",
            "xref_key": "glycan_xref_sandbox",
            "xref_id": "G00855BY",
            "protein_name": "Dol-P-Man:Man(5)GlcNAc(2)-PP-Dol alpha-1,3-mannosyltransferase",
            "gene": "ALG3",
            "gene_link": "https://www.uniprot.org/uniprotkb/Q92685",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human",
            "evidence": [
                {
                    "id": "G00855BY",
                    "database": "SandBox",
                    "url": "https://sandbox.glyomics.org/explore.html?G00855BY"
                }
            ]
        }
    ],
    "crossref": [
        {
            "id": "177784063",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/177784063",
            "database": "PubChem Compound",
            "categories": [
                "Chemical"
            ]
        },
        {
            "id": "521706603",
            "url": "https://pubchem.ncbi.nlm.nih.gov/substance/521706603",
            "database": "PubChem Substance",
            "categories": [
                "Chemical"
            ]
        },
        {
            "id": "G00855BY",
            "url": "https://glytoucan.org/Structures/Glycans/G00855BY",
            "database": "GlyTouCan",
            "categories": [
                "Glycomics"
            ]
        },
        {
            "id": "G00855BY",
            "url": "https://sandbox.glyomics.org/explore.html?G00855BY",
            "database": "SandBox",
            "categories": [
                "Enzyme and Pathway"
            ]
        },
        {
            "id": "G00855BY",
            "url": "https://gnome.glyomics.org/restrictions/GlyGen.StructureBrowser.html?focus=G00855BY",
            "database": "GNOme",
            "categories": [
                "Glycomics",
                "Ontology and Dictionary"
            ]
        },
        {
            "id": "G00855BY",
            "url": "https://glycosmos.org/glycans/show/G00855BY",
            "database": "GlyCosmos",
            "categories": [
                "Glycoproteomics",
                "Glycomics"
            ]
        }
    ],
    "glycoprotein": [
        {
            "uniprot_canonical_ac": "P08236-1",
            "start_pos": 173,
            "end_pos": 173,
            "residue": "Asn",
            "evidence": [
                {
                    "database": "PubMed",
                    "id": "37639587",
                    "url": "https://glygen.org/publication/PubMed/37639587"
                },
                {
                    "database": "TableMaker",
                    "id": "TP6077383",
                    "url": "https://glygen.ccrc.uga.edu/tablemaker/data/dataset/TP6077383"
                }
            ],
            "protein_name": "Beta-glucuronidase",
            "gene_name": "GUSB",
            "tax_id": 9606,
            "tax_name": "Homo sapiens",
            "tax_common_name": "Human"
        }
    ],
    "publication": [
        {
            "title": "Glycoproteome remodeling and organelle-specific N-glycosylation accompany neutrophil granulopoiesis.",
            "journal": "Proceedings of the National Academy of Sciences of the United States of America",
            "date": "2023-09-05",
            "authors": "Kawahara Rebeca, Ugonotti Julian, Chatterjee Sayantani, Tjondro Harry C, Loke Ian, Parker Benjamin L, Venkatakrishnan Vignesh, Dieckmann Regis, Sumer-Bayraktar Zeynep, Karlsson-Bengtsson Anna, Bylund Johan, Thaysen-Andersen Morten",
            "evidence": [
                {
                    "database": "TableMaker",
                    "id": "TP6077383",
                    "url": "https://glygen.ccrc.uga.edu/tablemaker/data/dataset/TP6077383"
                }
            ],
            "reference": [
                {
                    "type": "PubMed",
                    "id": "37639587",
                    "url": "https://glygen.org/publication/PubMed/37639587"
                }
            ]
        }
    ],
    "residues": [
        {
            "id": "NA",
            "name": "N-glycan_core_D-GlcNAc_A",
            "attachedby": "rxn.P42867-1",
            "detachedby": "",
            "parentid": "0"
        },
        {
            "id": "NA",
            "name": "N-glycan_core_D-GlcNAc_A",
            "attachedby": "rxn.Q9H3H5-1",
            "detachedby": "",
            "parentid": "0"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q8BML3-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q96F25-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q9D081-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q9D8C3-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NB",
            "name": "N-glycan_core_b-D-GlcpNAc_B",
            "attachedby": "rxn.Q9NP73-1",
            "detachedby": "",
            "parentid": "NA"
        },
        {
            "id": "NC",
            "name": "N-glycan_core_b-D-Manp_C",
            "attachedby": "rxn.Q921Q3-1",
            "detachedby": "",
            "parentid": "NB"
        },
        {
            "id": "NC",
            "name": "N-glycan_core_b-D-Manp_C",
            "attachedby": "rxn.Q9BT22-1",
            "detachedby": "",
            "parentid": "NB"
        },
        {
            "id": "N1",
            "name": "N-glycan_a-D-Manp_1",
            "attachedby": "rxn.Q9DBE8-1",
            "detachedby": "",
            "parentid": "NC"
        },
        {
            "id": "N1",
            "name": "N-glycan_a-D-Manp_1",
            "attachedby": "rxn.Q9H553-1",
            "detachedby": "",
            "parentid": "NC"
        },
        {
            "id": "N4",
            "name": "N-glycan_a-D-Manp_4",
            "attachedby": "rxn.Q9DBE8-1",
            "detachedby": "",
            "parentid": "NC"
        },
        {
            "id": "N4",
            "name": "N-glycan_a-D-Manp_4",
            "attachedby": "rxn.Q9H553-1",
            "detachedby": "",
            "parentid": "NC"
        },
        {
            "id": "N19",
            "name": "N-glycan_a-D-Manp_19",
            "attachedby": "rxn.Q8K2A8-1",
            "detachedby": "",
            "parentid": "N4"
        },
        {
            "id": "N19",
            "name": "N-glycan_a-D-Manp_19",
            "attachedby": "rxn.Q92685-1",
            "detachedby": "",
            "parentid": "N4"
        },
        {
            "id": "N20",
            "name": "N-glycan_a-D-Manp_20",
            "attachedby": "rxn.Q8K2A8-1",
            "detachedby": "",
            "parentid": "N4"
        },
        {
            "id": "N20",
            "name": "N-glycan_a-D-Manp_20",
            "attachedby": "rxn.Q92685-1",
            "detachedby": "",
            "parentid": "N4"
        }
    ],
    "composition": [
        {
            "name": "6-Deoxy-Hexose",
            "residue": "dhex",
            "count": 0,
            "cid": "840",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840"
        },
        {
            "name": "Hexose",
            "residue": "hex",
            "count": 9,
            "cid": "206",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206"
        },
        {
            "name": "N-Acetylhexosamine",
            "residue": "hexnac",
            "count": 2,
            "cid": "899",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899"
        },
        {
            "name": "Hexuronic Acid",
            "residue": "hexa",
            "count": 0,
            "cid": "610",
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610"
        },
        {
            "name": "2-Amino-2-Deoxy-Hexose",
            "residue": "hexn",
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