{
    "glycoprotein": [], 
    "number_monosaccharides": 21, 
    "inchi_key": {}, 
    "classification": [
        {
            "subtype": {
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=complex_N-glycan&jump_to_nav=true", 
                "name": "Complex"
            }, 
            "type": {
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=N-glycan&jump_to_nav=true", 
                "name": "N-linked"
            }
        }, 
        {
            "subtype": {
                "name": "Core-fucosylated"
            }, 
            "type": {
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=N-glycan&jump_to_nav=true", 
                "name": "N-linked"
            }
        }
    ], 
    "mass_pme": 4899.47, 
    "names": [], 
    "fully_determined": "", 
    "keywords": [], 
    "wurcs": "WURCS=2.0/7,21,20/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2122h-1x_1-5_2*NCC/3=O][a1221m-1a_1-5][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O]/1-1-2-3-4-5-6-7-3-4-5-6-1-5-6-7-4-5-6-7-5/a4-b1_a6-u1_b4-c1_c3-d1_c6-i1_e3-f1_e4-g1_g3-h2_j3-k1_j4-l1_l3-m1_m3-n1_m4-o1_o3-p2_q3-r1_q4-s1_s3-t2_d?-e1_i?-j1_i?-q1", 
    "species": [], 
    "publication": [
        {
            "reference": [
                {
                    "url": "https://www.ncbi.nlm.nih.gov/pubmed/24698316", 
                    "type": [
                        "PubMed"
                    ], 
                    "id": "24698316"
                }
            ], 
            "title": "Hypomorphic homozygous mutations in phosphoglucomutase 3 (PGM3) impair immunity and increase serum IgE levels.", 
            "journal": "The Journal of allergy and clinical immunology", 
            "evidence": [
                {
                    "url": "http://www.unicarbkb.org/structure/8155", 
                    "id": "8155", 
                    "database": "UniCarbKB"
                }
            ], 
            "authors": "Sassi A, Lazaroski S, Wu G, Haslam SM, Fliegauf M, Mellouli F, Patiroglu T, Unal E, Ozdemir MA, Jouhadi Z, Khadir K, Ben-Khemis L, Ben-Ali M, Ben-Mustapha I, Borchani L, Pfeifer D, Jakob T, Khemiri M, Asplund AC, Gustafsson MO, Lundin KE, Falk-Sorqvist E, Moens LN, Gungor HE, Engelhardt KR, Dziadzio M, Stauss H, Fleckenstein B, Meier R, Prayitno K, Maul-Pavicic A, Schaffer S, Rakhmanov M, Henneke P, Kraus H, Eibel H, Kolsch U, Nadifi S, Nilsson M, Bejaoui M, Schaffer AA, Smith CI, Dell A, Barbouche MR, Grimbacher B", 
            "date": "2014"
        }, 
        {
            "reference": [
                {
                    "url": "https://www.ncbi.nlm.nih.gov/pubmed/21852454", 
                    "type": [
                        "PubMed"
                    ], 
                    "id": "21852454"
                }
            ], 
            "title": "Human sperm binding is mediated by the sialyl-Lewis(x) oligosaccharide on the zona pellucida.", 
            "journal": "Science (New York, N.Y.)", 
            "evidence": [
                {
                    "url": "http://www.unicarbkb.org/structure/8155", 
                    "id": "8155", 
                    "database": "UniCarbKB"
                }
            ], 
            "authors": "Pang PC, Chiu PC, Lee CL, Chang LY, Panico M, Morris HR, Haslam SM, Khoo KH, Clark GF, Yeung WS, Dell A", 
            "date": "2011"
        }, 
        {
            "reference": [
                {
                    "url": "https://www.ncbi.nlm.nih.gov/pubmed/23829188", 
                    "type": [
                        "PubMed"
                    ], 
                    "id": "23829188"
                }
            ], 
            "title": "N-glycosylation profile of undifferentiated and adipogenically differentiated human bone marrow mesenchymal stem cells: towards a next generation of stem cell markers.", 
            "journal": "Stem cells and development", 
            "evidence": [
                {
                    "url": "http://www.unicarbkb.org/structure/8155", 
                    "id": "8155", 
                    "database": "UniCarbKB"
                }
            ], 
            "authors": "Hamouda H, Ullah M, Berger M, Sittinger M, Tauber R, Ringe J, Blanchard V", 
            "date": "2013"
        }
    ], 
    "subsumption": [
        {
            "type": "basecomposition", 
            "id": "G53622FH", 
            "relationship": "basecomposition"
        }, 
        {
            "type": "composition", 
            "id": "G08324RM", 
            "relationship": "composition"
        }
    ], 
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q9D081-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14 homolog", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Alg14", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "P42867-1", 
            "gene_link": "https://www.uniprot.org/uniprot/P42867", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Dpagt1", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9WTS2-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9WTS2", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-(1,6)-fucosyltransferase", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Fut8", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9H553-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ALG2", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9H3H5-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "DPAGT1", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9BYC5-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9BYC5", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-(1,6)-fucosyltransferase", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "FUT8", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9BT22-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ALG1", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q8BML3-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "N-acetylglucosaminyldiphosphodolichol N-acetylglucosaminyltransferase", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Glt28d2", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q96F25-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14 homolog", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ALG14", 
            "tax_id": 9606
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        {
            "uniprot_canonical_ac": "Q9NP73-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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            ], 
            "protein_name": "Putative bifunctional UDP-N-acetylglucosamine transferase and deubiquitinase ALG13", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ALG13", 
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        }, 
        {
            "uniprot_canonical_ac": "Q9D8C3-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3", 
            "evidence": [
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                    "url": "https://data.glygen.org/GLY_000284", 
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                }
            ], 
            "protein_name": "Putative bifunctional UDP-N-acetylglucosamine transferase and deubiquitinase ALG13", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
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            "tax_id": 10090
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        {
            "uniprot_canonical_ac": "Q921Q3-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Alg1", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9DBE8-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Alg2", 
            "tax_id": 10090
        }
    ], 
    "motifs": [
        {
            "synonym": "", 
            "id": "GGM.000034", 
            "name": "3-Sialyl-LN (type 2)"
        }, 
        {
            "synonym": "SSEA-1; Lex; Le^x", 
            "id": "GGM.000018", 
            "name": "Lewis x"
        }, 
        {
            "synonym": "", 
            "id": "GGM.000052", 
            "name": "Polysialic acid"
        }, 
        {
            "synonym": "Slex; SLex; SLe^x", 
            "id": "GGM.000022", 
            "name": "Sialyl Lewis x"
        }, 
        {
            "synonym": "N-acetyllactosamine", 
            "id": "GGM.000001", 
            "name": "Type 2 LN"
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    ], 
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    "composition": [
        {
            "residue": "p", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061", 
            "name": "Phosphate", 
            "cid": "1061"
        }, 
        {
            "residue": "hex", 
            "count": 7, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206", 
            "name": "Hexose", 
            "cid": "206"
        }, 
        {
            "residue": "hexn", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739", 
            "name": "2-Amino-2-Deoxy-Hexose", 
            "cid": "739"
        }, 
        {
            "residue": "dhex", 
            "count": 5, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840", 
            "name": "6-Deoxy-Hexose", 
            "cid": "840"
        }, 
        {
            "residue": "hexa", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610", 
            "name": "Hexuronic Acid", 
            "cid": "610"
        }, 
        {
            "residue": "other", 
            "count": 0, 
            "name": "Other"
        }, 
        {
            "residue": "s", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117", 
            "name": "Sulfate", 
            "cid": "1117"
        }, 
        {
            "residue": "neuac", 
            "count": 3, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197", 
            "name": "N-Acetyl-Neuraminic Acid", 
            "cid": "439197"
        }, 
        {
            "residue": "hexnac", 
            "count": 6, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899", 
            "name": "N-Acetylhexosamine", 
            "cid": "899"
        }, 
        {
            "residue": "neugc", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001", 
            "name": "N-Glycolyl-Neuraminic Acid", 
            "cid": "440001"
        }
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    "interactions": [], 
    "residues": [
        {
            "attachedby": "rxn.Q96F25-1", 
            "detachedby": "", 
            "id": "NB", 
            "parentid": "NA", 
            "name": "N-glycan_core_b-D-GlcpNAc_B"
        }, 
        {
            "attachedby": "rxn.Q9D081-1", 
            "detachedby": "", 
            "id": "NB", 
            "parentid": "NA", 
            "name": "N-glycan_core_b-D-GlcpNAc_B"
        }, 
        {
            "attachedby": "rxn.Q9DBE8-1", 
            "detachedby": "", 
            "id": "N1", 
            "parentid": "NC", 
            "name": "N-glycan_a-D-Manp_1"
        }, 
        {
            "attachedby": "rxn.Q9H553-1", 
            "detachedby": "", 
            "id": "N1", 
            "parentid": "NC", 
            "name": "N-glycan_a-D-Manp_1"
        }, 
        {
            "attachedby": "rxn.Q9DBE8-1", 
            "detachedby": "", 
            "id": "N4", 
            "parentid": "NC", 
            "name": "N-glycan_a-D-Manp_4"
        }, 
        {
            "attachedby": "rxn.Q9H553-1", 
            "detachedby": "", 
            "id": "N4", 
            "parentid": "NC", 
            "name": "N-glycan_a-D-Manp_4"
        }, 
        {
            "attachedby": "rxn.Q9BYC5-1", 
            "detachedby": "", 
            "id": "N15", 
            "parentid": "NA", 
            "name": "N-glycan_a-L-Fucp_15"
        }, 
        {
            "attachedby": "rxn.Q9WTS2-1", 
            "detachedby": "", 
            "id": "N15", 
            "parentid": "NA", 
            "name": "N-glycan_a-L-Fucp_15"
        }, 
        {
            "attachedby": "rxn.Q8BML3-1", 
            "detachedby": "", 
            "id": "NB", 
            "parentid": "NA", 
            "name": "N-glycan_core_b-D-GlcpNAc_B"
        }, 
        {
            "attachedby": "rxn.Q921Q3-1", 
            "detachedby": "", 
            "id": "NC", 
            "parentid": "NB", 
            "name": "N-glycan_core_b-D-Manp_C"
        }, 
        {
            "attachedby": "rxn.Q9BT22-1", 
            "detachedby": "", 
            "id": "NC", 
            "parentid": "NB", 
            "name": "N-glycan_core_b-D-Manp_C"
        }, 
        {
            "attachedby": "rxn.Q9D8C3-1", 
            "detachedby": "", 
            "id": "NB", 
            "parentid": "NA", 
            "name": "N-glycan_core_b-D-GlcpNAc_B"
        }, 
        {
            "attachedby": "rxn.Q9NP73-1", 
            "detachedby": "", 
            "id": "NB", 
            "parentid": "NA", 
            "name": "N-glycan_core_b-D-GlcpNAc_B"
        }, 
        {
            "attachedby": "rxn.P42867-1", 
            "detachedby": "", 
            "id": "NA", 
            "parentid": "0", 
            "name": "N-glycan_core_D-GlcNAc_A"
        }, 
        {
            "attachedby": "rxn.Q9H3H5-1", 
            "detachedby": "", 
            "id": "NA", 
            "parentid": "0", 
            "name": "N-glycan_core_D-GlcNAc_A"
        }
    ], 
    "glycam": "", 
    "inchi": "", 
    "crossref": [
        {
            "url": "http://www.unicarbkb.org/structure/8155", 
            "id": "8155", 
            "database": "UniCarbKB"
        }, 
        {
            "url": "https://glytoucan.org/Structures/Glycans/G09405BO", 
            "id": "G09405BO", 
            "database": "GlyTouCan"
        }, 
        {
            "url": "https://www.glygen.org/glycan/G09405BO", 
            "id": "G09405BO", 
            "database": "GNOme"
        }, 
        {
            "url": "https://raw.githack.com/CCRC-will/glycoTree/master/portal/explore.html?G09405BO", 
            "id": "G09405BO", 
            "database": "SandBox"
        }
    ], 
    "tool_support": {
        "sandbox": "yes", 
        "gnome": "yes"
    }, 
    "glycoct": "RES 1b:b-dglc-HEX-1:5 2s:n-acetyl 3b:b-dglc-HEX-1:5 4s:n-acetyl 5b:b-dman-HEX-1:5 6b:a-dman-HEX-1:5 7b:x-dglc-HEX-1:5 8s:n-acetyl 9b:a-lgal-HEX-1:5|6:d 10b:b-dgal-HEX-1:5 11b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 12s:n-acetyl 13b:a-dman-HEX-1:5 14b:x-dglc-HEX-1:5 15s:n-acetyl 16b:a-lgal-HEX-1:5|6:d 17b:b-dgal-HEX-1:5 18b:b-dglc-HEX-1:5 19s:n-acetyl 20b:a-lgal-HEX-1:5|6:d 21b:b-dgal-HEX-1:5 22b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 23s:n-acetyl 24b:x-dglc-HEX-1:5 25s:n-acetyl 26b:a-lgal-HEX-1:5|6:d 27b:b-dgal-HEX-1:5 28b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 29s:n-acetyl 30b:a-lgal-HEX-1:5|6:d LIN 1:1d(2+1)2n 2:1o(4+1)3d 3:3d(2+1)4n 4:3o(4+1)5d 5:5o(3+1)6d 6:6o(-1+1)7d 7:7d(2+1)8n 8:7o(3+1)9d 9:7o(4+1)10d 10:10o(3+2)11d 11:11d(5+1)12n 12:5o(6+1)13d 13:13o(-1+1)14d 14:14d(2+1)15n 15:14o(3+1)16d 16:14o(4+1)17d 17:17o(3+1)18d 18:18d(2+1)19n 19:18o(3+1)20d 20:18o(4+1)21d 21:21o(3+2)22d 22:22d(5+1)23n 23:13o(-1+1)24d 24:24d(2+1)25n 25:24o(3+1)26d 26:24o(4+1)27d 27:27o(3+2)28d 28:28d(5+1)29n 29:1o(6+1)30d", 
    "iupac": "alpha-D-NeupAc-(2->3)-beta-D-Galp-(1->4)[alpha-L-Fucp-(1->3)]-?-D-GlcpNAc-(1->?)-alpha-D-Manp-(1->3)[alpha-D-NeupAc-(2->3)-beta-D-Galp-(1->4)[alpha-L-Fucp-(1->3)]-beta-D-GlcpNAc-(1->3)-beta-D-Galp-(1->4)[alpha-L-Fucp-(1->3)]-?-D-GlcpNAc-(1->?)[alpha-D-NeupAc-(2->3)-beta-D-Galp-(1->4)[alpha-L-Fucp-(1->3)]-?-D-GlcpNAc-(1->?)]-alpha-D-Manp-(1->6)]-beta-D-Manp-(1->4)-beta-D-GlcpNAc-(1->4)[alpha-L-Fucp-(1->6)]-beta-D-GlcpNAc-(1->", 
    "smiles_isomeric": "", 
    "mass": 3974.43, 
    "expression": []
}