{
    "glycoprotein": [
        {
            "uniprot_canonical_ac": "P14780-1", 
            "protein_name": "Matrix metalloproteinase-9", 
            "tax_name": "Homo sapiens", 
            "tax_id": 9606, 
            "gene_name": "MMP9", 
            "evidence": [
                {
                    "url": "https://glyconnect.expasy.org/browser/proteins/6", 
                    "id": "6", 
                    "database": "GlyConnect"
                }, 
                {
                    "url": "https://www.ncbi.nlm.nih.gov/pubmed/11123894", 
                    "id": "11123894", 
                    "database": "PubMed"
                }
            ]
        }
    ], 
    "number_monosaccharides": 8, 
    "inchi_key": {}, 
    "classification": [
        {
            "subtype": {
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=core-2_O-glycan&jump_to_nav=true", 
                "name": "Core 2"
            }, 
            "type": {
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=O-glycan&jump_to_nav=true", 
                "name": "O-linked"
            }
        }
    ], 
    "mass_pme": 2116.07, 
    "names": [], 
    "fully_determined": "", 
    "keywords": [], 
    "wurcs": "WURCS=2.0/4,8,7/[a2112h-1a_1-5_2*NCC/3=O][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O][a2122h-1b_1-5_2*NCC/3=O]/1-2-3-4-2-4-2-3/a3-b1_a6-d1_b3-c2_d4-e1_f4-g1_g6-h2_e?-f1", 
    "species": [
        {
            "common_name": "Human", 
            "annotation_category": "Direct", 
            "evidence": [
                {
                    "url": "https://glyconnect.expasy.org/browser/structures/2841", 
                    "id": "2841", 
                    "database": "GlyConnect"
                }
            ], 
            "taxid": 9606, 
            "name": "Homo sapiens"
        }
    ], 
    "publication": [
        {
            "reference": [
                {
                    "url": "https://www.ncbi.nlm.nih.gov/pubmed/11123894", 
                    "type": [
                        "PubMed"
                    ], 
                    "id": "11123894"
                }
            ], 
            "title": "O-glycan analysis of natural human neutrophil gelatinase B using a combination of normal phase-HPLC and online tandem mass spectrometry: implications for the domain organization of the enzyme.", 
            "journal": "Biochemistry", 
            "evidence": [
                {
                    "url": "https://glyconnect.expasy.org/browser/proteins/6", 
                    "id": "6", 
                    "database": "GlyConnect"
                }
            ], 
            "authors": "Mattu TS, Royle L, Langridge J, Wormald MR, Van den Steen PE, Van Damme J, Opdenakker G, Harvey DJ, Dwek RA, Rudd PM", 
            "date": "2000"
        }
    ], 
    "subsumption": [
        {
            "type": "basecomposition", 
            "id": "G13058MK", 
            "relationship": "basecomposition"
        }, 
        {
            "type": "composition", 
            "id": "G55239YU", 
            "relationship": "composition"
        }, 
        {
            "type": "topology", 
            "id": "G67293HE", 
            "relationship": "topology"
        }
    ], 
    "enzyme": [], 
    "motifs": [
        {
            "synonym": "N-acetyllactosamine", 
            "id": "GGM.000001", 
            "name": "Type 2 LN"
        }
    ], 
    "glytoucan_ac": "G05380VG", 
    "composition": [
        {
            "residue": "hexa", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610", 
            "name": "Hexuronic Acid", 
            "cid": "610"
        }, 
        {
            "residue": "hexnac", 
            "count": 3, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899", 
            "name": "N-Acetylhexosamine", 
            "cid": "899"
        }, 
        {
            "residue": "neuac", 
            "count": 2, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197", 
            "name": "N-Acetyl-Neuraminic Acid", 
            "cid": "439197"
        }, 
        {
            "residue": "neugc", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001", 
            "name": "N-Glycolyl-Neuraminic Acid", 
            "cid": "440001"
        }, 
        {
            "residue": "s", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117", 
            "name": "Sulfate", 
            "cid": "1117"
        }, 
        {
            "residue": "other", 
            "count": 0, 
            "name": "Other"
        }, 
        {
            "residue": "hex", 
            "count": 3, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206", 
            "name": "Hexose", 
            "cid": "206"
        }, 
        {
            "residue": "p", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061", 
            "name": "Phosphate", 
            "cid": "1061"
        }, 
        {
            "residue": "dhex", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840", 
            "name": "6-Deoxy-Hexose", 
            "cid": "840"
        }, 
        {
            "residue": "hexn", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739", 
            "name": "2-Amino-2-Deoxy-Hexose", 
            "cid": "739"
        }
    ], 
    "interactions": [], 
    "residues": [], 
    "glycam": "", 
    "inchi": "", 
    "crossref": [
        {
            "url": "https://glyconnect.expasy.org/browser/structures/2841", 
            "id": "2841", 
            "database": "GlyConnect"
        }, 
        {
            "url": "https://www.glygen.org/glycan/G05380VG", 
            "id": "G05380VG", 
            "database": "GNOme"
        }, 
        {
            "url": "https://glytoucan.org/Structures/Glycans/G05380VG", 
            "id": "G05380VG", 
            "database": "GlyTouCan"
        }
    ], 
    "tool_support": {
        "sandbox": "no", 
        "gnome": "yes"
    }, 
    "glycoct": "RES 1b:a-dgal-HEX-1:5 2s:n-acetyl 3b:b-dgal-HEX-1:5 4b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 5s:n-acetyl 6b:b-dglc-HEX-1:5 7s:n-acetyl 8b:b-dgal-HEX-1:5 9b:b-dglc-HEX-1:5 10s:n-acetyl 11b:b-dgal-HEX-1:5 12b:a-dgro-dgal-NON-2:6|1:a|2:keto|3:d 13s:n-acetyl LIN 1:1d(2+1)2n 2:1o(3+1)3d 3:3o(3+2)4d 4:4d(5+1)5n 5:1o(6+1)6d 6:6d(2+1)7n 7:6o(4+1)8d 8:8o(-1+1)9d 9:9d(2+1)10n 10:9o(4+1)11d 11:11o(6+2)12d 12:12d(5+1)13n", 
    "iupac": "alpha-D-NeupAc-(2->3)-beta-D-Galp-(1->3)[alpha-D-NeupAc-(2->6)-beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->?)-beta-D-Galp-(1->4)-beta-D-GlcpNAc-(1->6)]-alpha-D-GalpNAc-(1->", 
    "smiles_isomeric": "", 
    "mass": 1695.6, 
    "expression": []
}