{
    "glycoprotein": [], 
    "number_monosaccharides": 13, 
    "inchi_key": {
        "url": "https://pubchem.ncbi.nlm.nih.gov/compound/OKIGEKOBSZKGOM-JGHNGCCPSA-N", 
        "key": "OKIGEKOBSZKGOM-JGHNGCCPSA-N"
    }, 
    "classification": [
        {
            "subtype": {
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=complex_N-glycan&jump_to_nav=true", 
                "name": "Complex"
            }, 
            "type": {
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=N-glycan&jump_to_nav=true", 
                "name": "N-linked"
            }
        }
    ], 
    "mass_pme": 3218.62, 
    "names": [], 
    "fully_determined": "yes", 
    "keywords": [
        "fully_determined"
    ], 
    "wurcs": "WURCS=2.0/5,13,12/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a2112h-1b_1-5][Aad21122h-2a_2-6_5*NCC/3=O]/1-1-2-3-1-4-1-4-5-3-1-4-5/a4-b1_b4-c1_c3-d1_c6-j1_d2-e1_d4-g1_e4-f1_g4-h1_h3-i2_j2-k1_k4-l1_l6-m2", 
    "species": [], 
    "publication": [], 
    "subsumption": [
        {
            "type": "topology", 
            "id": "G08875HF", 
            "relationship": "topology"
        }, 
        {
            "type": "composition", 
            "id": "G60942FK", 
            "relationship": "composition"
        }, 
        {
            "type": "basecomposition", 
            "id": "G34447YB", 
            "relationship": "basecomposition"
        }
    ], 
    "enzyme": [
        {
            "uniprot_canonical_ac": "O60909-1", 
            "gene_link": "https://www.uniprot.org/uniprot/O60909", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Beta-1,4-galactosyltransferase 2", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "B4GALT2", 
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        {
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            "gene_link": "https://www.uniprot.org/uniprot/Q10469", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
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            ], 
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            "gene": "MGAT2", 
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        {
            "uniprot_canonical_ac": "Q96F25-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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                }
            ], 
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            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ALG14", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9D081-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
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                }
            ], 
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14 homolog", 
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            "tax_common_name": "Mouse", 
            "gene": "Alg14", 
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        }, 
        {
            "uniprot_canonical_ac": "P42867-1", 
            "gene_link": "https://www.uniprot.org/uniprot/P42867", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
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            ], 
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Dpagt1", 
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        {
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            "gene_link": "https://www.uniprot.org/uniprot/Q8VIB3", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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                }
            ], 
            "protein_name": "Type 2 lactosamine alpha-2,3-sialyltransferase", 
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            "gene": "St3gal6", 
            "tax_id": 10090
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        {
            "uniprot_canonical_ac": "P97325-1", 
            "gene_link": "https://www.uniprot.org/uniprot/P97325", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "CMP-N-acetylneuraminate-beta-1,4-galactoside alpha-2,3-sialyltransferase", 
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            "gene": "St3gal3", 
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        }, 
        {
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            "gene_link": "https://www.uniprot.org/uniprot/Q9H553", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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                }
            ], 
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ALG2", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9H3H5-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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                }
            ], 
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase", 
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            "gene": "DPAGT1", 
            "tax_id": 9606
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        {
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            "gene_link": "https://www.uniprot.org/uniprot/Q921V5", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-1,6-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Mgat2", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q64685-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q64685", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 1", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "St6gal1", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "P15291-1", 
            "gene_link": "https://www.uniprot.org/uniprot/P15291", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Processed beta-1,4-galactosyltransferase 1", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "B4GALT1", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9UM21-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9UM21", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase A", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "MGAT4A", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q76K27-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q76K27", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 2", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "St6gal2", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9D306-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9D306", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
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            ], 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase C", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Mgat4c", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q921Q3-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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                }
            ], 
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Alg1", 
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        }, 
        {
            "uniprot_canonical_ac": "Q812F8-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q812F8", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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            ], 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase B", 
            "tax_name": "Mus musculus", 
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        {
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            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73", 
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                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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            ], 
            "protein_name": "Putative bifunctional UDP-N-acetylglucosamine transferase and deubiquitinase ALG13", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ALG13", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9UQ53-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9UQ53", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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                }
            ], 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase B", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "MGAT4B", 
            "tax_id": 9606
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        {
            "uniprot_canonical_ac": "Q9Y274-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9Y274", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
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            ], 
            "protein_name": "Type 2 lactosamine alpha-2,3-sialyltransferase", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ST3GAL6", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q96JF0-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q96JF0", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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                }
            ], 
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 2", 
            "tax_name": "Homo sapiens", 
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        {
            "uniprot_canonical_ac": "Q812G0-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q812G0", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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                }
            ], 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase A", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Mgat4a", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9D8C3-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
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                }
            ], 
            "protein_name": "Putative bifunctional UDP-N-acetylglucosamine transferase and deubiquitinase ALG13", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Alg13", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "P15535-1", 
            "gene_link": "https://www.uniprot.org/uniprot/P15535", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Processed beta-1,4-galactosyltransferase 1", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "B4galt1", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9UBM8-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9UBM8", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase C", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "MGAT4C", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q11206-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q11206", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 4", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ST3GAL4", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "P27808-1", 
            "gene_link": "https://www.uniprot.org/uniprot/P27808", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Mgat1", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9DBE8-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Alg2", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q11203-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q11203", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "CMP-N-acetylneuraminate-beta-1,4-galactoside alpha-2,3-sialyltransferase", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ST3GAL3", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "O60512-1", 
            "gene_link": "https://www.uniprot.org/uniprot/O60512", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Beta-1,4-galactosyltransferase 3", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "B4GALT3", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "P15907-1", 
            "gene_link": "https://www.uniprot.org/uniprot/P15907", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Beta-galactoside alpha-2,6-sialyltransferase 1", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ST6GAL1", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9BT22-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "ALG1", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q91Y74-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q91Y74", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 4", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "St3gal4", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "P26572-1", 
            "gene_link": "https://www.uniprot.org/uniprot/P26572", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 2-beta-N-acetylglucosaminyltransferase", 
            "tax_name": "Homo sapiens", 
            "tax_common_name": "Human", 
            "gene": "MGAT1", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q8BML3-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "N-acetylglucosaminyldiphosphodolichol N-acetylglucosaminyltransferase", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "Glt28d2", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q91YY2-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q91YY2", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Beta-1,4-galactosyltransferase 3", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "B4galt3", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9Z2Y2-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9Z2Y2", 
            "evidence": [
                {
                    "url": "https://data.glygen.org/GLY_000284", 
                    "id": "GLY_000284", 
                    "database": "GlyGen"
                }
            ], 
            "protein_name": "Beta-1,4-galactosyltransferase 2", 
            "tax_name": "Mus musculus", 
            "tax_common_name": "Mouse", 
            "gene": "B4galt2", 
            "tax_id": 10090
        }
    ], 
    "motifs": [
        {
            "synonym": "", 
            "id": "GGM.000034", 
            "name": "3-Sialyl-LN (type 2)"
        }, 
        {
            "synonym": "", 
            "id": "GGM.000052", 
            "name": "Polysialic acid"
        }, 
        {
            "synonym": "N-acetyllactosamine", 
            "id": "GGM.000001", 
            "name": "Type 2 LN"
        }
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    "crossref": [
        {
            "url": "http://www.glycosciences.de/database/start.php?action=explore_linucsid&linucsid=24867", 
            "id": "24867", 
            "database": "Glycosciences.de"
        }, 
        {
            "url": "https://www.glygen.org/glycan/G03385XG", 
            "id": "G03385XG", 
            "database": "GNOme"
        }, 
        {
            "url": "https://raw.githack.com/CCRC-will/glycoTree/master/portal/explore.html?G03385XG", 
            "id": "G03385XG", 
            "database": "SandBox"
        }, 
        {
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            "id": "252286000", 
            "database": "PubChem Substance"
        }, 
        {
            "url": "https://glytoucan.org/Structures/Glycans/G03385XG", 
            "id": "G03385XG", 
            "database": "GlyTouCan"
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        {
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            "id": "91853560", 
            "database": "PubChem Compound"
        }, 
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            "id": "146552", 
            "database": "ChEBI"
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    "tool_support": {
        "sandbox": "yes", 
        "gnome": "yes"
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    "mass": 2587.92, 
    "expression": []
}