{
    "glytoucan_ac": "G11994QC", 
    "mass": 1259.47, 
    "iupac": "beta-D-GlcpNAc-(1->4)-alpha-D-Manp-(1->3)[alpha-D-Manp-(1->6)]-beta-D-Manp-(1->4)-beta-D-GlcpNAc-(1->4)[alpha-L-Fucp-(1->6)]-beta-D-GlcpNAc(1->", 
    "wurcs": "WURCS=2.0/4,7,6/[a2122h-1b_1-5_2*NCC/3=O][a1122h-1b_1-5][a1122h-1a_1-5][a1221m-1a_1-5]/1-1-2-3-1-3-4/a4-b1_a6-g1_b4-c1_c3-d1_c6-f1_d4-e1", 
    "glycoct": "RES\\n1b:b-dglc-HEX-1:5\\n2s:n-acetyl\\n3b:b-dglc-HEX-1:5\\n4s:n-acetyl\\n5b:b-dman-HEX-1:5\\n6b:a-dman-HEX-1:5\\n7b:b-dglc-HEX-1:5\\n8s:n-acetyl\\n9b:a-dman-HEX-1:5\\n10b:a-lgal-HEX-1:5|6:d\\nLIN\\n1:1d(2+1)2n\\n2:1o(4+1)3d\\n3:3d(2+1)4n\\n4:3o(4+1)5d\\n5:5o(3+1)6d\\n6:6o(4+1)7d\\n7:7d(2+1)8n\\n8:5o(6+1)9d\\n9:1o(6+1)10d", 
    "species": [
        {
            "evidence": [
                {
                    "url": "http://www.unicarbkb.org/structure/4211", 
                    "id": "4211", 
                    "database": "UniCarbKB"
                }
            ], 
            "taxid": 9606, 
            "name": "Homo sapiens"
        }
    ], 
    "classification": [
        {
            "subtype": {
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=Other&jump_to_nav=true", 
                "name": "Other"
            }, 
            "type": {
                "url": "http://bioportal.bioontology.org/ontologies/GLYCO/?p=classes&conceptid=N-glycan&jump_to_nav=true", 
                "name": "N-glycan"
            }
        }
    ], 
    "enzyme": [
        {
            "uniprot_canonical_ac": "Q921Q3-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q921Q3", 
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase; 2.4.1.142; Beta-1,4-mannosyltransferase; Asparagine-linked glycosylation protein 1 homolog; GDP-Man:GlcNAc2-PP-dolichol mannosyltransferase; GDP-mannose-dolichol diphosphochitobiose mannosyltransferase.", 
            "tax_name": "Mus musculus", 
            "gene": "Alg1; HMAT1; HMT1.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9D8C3-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9D8C3", 
            "protein_name": "Putative bifunctional UDP-N-acetylglucosamine transferase and deubiquitinase ALG13; 2.4.1.141; 3.4.19.12; Asparagine-linked glycosylation 13 homolog; Glycosyltransferase 28 domain-containing protein 1; UDP-N-acetylglucosamine transferase subunit ALG13 homolog.", 
            "tax_name": "Mus musculus", 
            "gene": "Alg13; Glt28d1; 2810046O15Rik; 4833435D08Rik; MDS031.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9D081-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9D081", 
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14 homolog.", 
            "tax_name": "Mus musculus", 
            "gene": "Alg14; 5430428G01Rik; AI854024.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9DBE8-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9DBE8", 
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2; 2.4.1.132; 2.4.1.257; GDP-Man:Man(2)GlcNAc(2)-PP-Dol alpha-1,6-mannosyltransferase; GDP-Man:Man(1)GlcNAc(2)-PP-Dol alpha-1,3-mannosyltransferase; Asparagine-linked glycosylation protein 2 homolog; GDP-Man:Man(1)GlcNAc(2)-PP-dolichol mannosyltransferase.", 
            "tax_name": "Mus musculus", 
            "gene": "Alg2; MNCb-5081; 1110018A23Rik; 1300013N08Rik; ALPG2; CDGIi.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "P42867-1", 
            "gene_link": "https://www.uniprot.org/uniprot/P42867", 
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase; 2.7.8.15; GlcNAc-1-P transferase; GPT; G1PT; N-acetylglucosamine-1-phosphate transferase.", 
            "tax_name": "Mus musculus", 
            "gene": "Dpagt1; Dpagt2; Gnpta; AU021132; G1PT; GPT.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9WTS2-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9WTS2", 
            "protein_name": "Alpha-(1,6)-fucosyltransferase; Alpha1-6FucT; 2.4.1.68; GDP-L-Fuc:N-acetyl-beta-D-glucosaminide alpha1,6-fucosyltransferase; Glycoprotein 6-alpha-L-fucosyltransferase; GDP-fucose--glycoprotein fucosyltransferase; Fucosyltransferase 8.", 
            "tax_name": "Mus musculus", 
            "gene": "Fut8.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q8BML3-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q8BML3", 
            "protein_name": "MCG145726; Glycosyltransferase 28 domain-containing 2.", 
            "tax_name": "Mus musculus", 
            "gene": "Glt28d2; mCG_145726; 4732486J07Rik; Glt28d1l.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q812G0-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q812G0", 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase A; 2.4.1.145; N-glycosyl-oligosaccharide-glycoprotein N-acetylglucosaminyltransferase IVa; GlcNAc-T IVa; GnT-IVa; N-acetylglucosaminyltransferase IVa; UDP-N-acetylglucosamine: alpha-1,3-D-mannoside beta-1,4-N-acetylglucosaminyltransferase IVa.", 
            "tax_name": "Mus musculus", 
            "gene": "Mgat4a; 9530018I07Rik; glcNAc-T-IVa; GnT-IVa.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q812F8-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q812F8", 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase B; 2.4.1.145; N-glycosyl-oligosaccharide-glycoprotein N-acetylglucosaminyltransferase IVb; GlcNAc-T IVb; GnT-IVb; N-acetylglucosaminyltransferase IVb; UDP-N-acetylglucosamine: alpha-1,3-D-mannoside beta-1,4-N-acetylglucosaminyltransferase IVb.", 
            "tax_name": "Mus musculus", 
            "gene": "Mgat4b; AA407995; GnTIVb.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9D306-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9D306", 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase C; 2.4.1.145; N-glycosyl-oligosaccharide-glycoprotein N-acetylglucosaminyltransferase IVc; GnT-IVc; N-acetylglucosaminyltransferase IVc; UDP-N-acetylglucosamine: alpha-1,3-D-mannoside beta-1,4-N-acetylglucosaminyltransferase IVc.", 
            "tax_name": "Mus musculus", 
            "gene": "Mgat4c; 9130411I17Rik; GntIVh.", 
            "tax_id": 10090
        }, 
        {
            "uniprot_canonical_ac": "Q9BT22-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9BT22", 
            "protein_name": "Chitobiosyldiphosphodolichol beta-mannosyltransferase; 2.4.1.142; Mannosyltransferase-1; MT-1; hMat-1; Beta-1,4-mannosyltransferase; Asparagine-linked glycosylation protein 1 homolog; GDP-Man:GlcNAc2-PP-dolichol mannosyltransferase; GDP-mannose-dolichol diphosphochitobiose mannosyltransferase.", 
            "tax_name": "Homo sapiens", 
            "gene": "ALG1; HMAT1; PSEC0061; HMT1; PRO1870; CDG1K; hMat-1; HMT-1; Mat-1; MT-1.", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9NP73-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9NP73", 
            "protein_name": "Putative bifunctional UDP-N-acetylglucosamine transferase and deubiquitinase ALG13; 2.4.1.141; 3.4.19.12; Asparagine-linked glycosylation 13 homolog; Glycosyltransferase 28 domain-containing protein 1; UDP-N-acetylglucosamine transferase subunit ALG13 homolog.", 
            "tax_name": "Homo sapiens", 
            "gene": "ALG13; CXorf45; MDS031; GLT28D1; CDG1S; EIEE36; TDRD13; YGL047W.", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q96F25-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q96F25", 
            "protein_name": "UDP-N-acetylglucosamine transferase subunit ALG14 homolog.", 
            "tax_name": "Homo sapiens", 
            "gene": "ALG14; CMS15.", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9H553-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9H553", 
            "protein_name": "Alpha-1,3/1,6-mannosyltransferase ALG2; 2.4.1.132; 2.4.1.257; GDP-Man:Man(2)GlcNAc(2)-PP-Dol alpha-1,6-mannosyltransferase; GDP-Man:Man(1)GlcNAc(2)-PP-Dol alpha-1,3-mannosyltransferase; Asparagine-linked glycosylation protein 2 homolog; GDP-Man:Man(1)GlcNAc(2)-PP-dolichol mannosyltransferase.", 
            "tax_name": "Homo sapiens", 
            "gene": "ALG2; PRO1298; CDG1I; CDGIi; CMS14; CMSTA3; hALPG2; NET38.", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9H3H5-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9H3H5", 
            "protein_name": "UDP-N-acetylglucosamine--dolichyl-phosphate N-acetylglucosaminephosphotransferase; 2.7.8.15; GlcNAc-1-P transferase; GPT; G1PT; N-acetylglucosamine-1-phosphate transferase.", 
            "tax_name": "Homo sapiens", 
            "gene": "DPAGT1; DPAGT2; ALG7; CDG-Ij; CDG1J; CMS13; CMSTA2; D11S366; DGPT; DPAGT; G1PT; GPT; UAGT; UGAT.", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9BYC5-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9BYC5", 
            "protein_name": "Alpha-(1,6)-fucosyltransferase; Alpha1-6FucT; 2.4.1.68; GDP-L-Fuc:N-acetyl-beta-D-glucosaminide alpha1,6-fucosyltransferase; Glycoprotein 6-alpha-L-fucosyltransferase; GDP-fucose--glycoprotein fucosyltransferase; Fucosyltransferase 8.", 
            "tax_name": "Homo sapiens", 
            "gene": "FUT8; CDGF; CDGF1.", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9UM21-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9UM21", 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase A; 2.4.1.145; N-glycosyl-oligosaccharide-glycoprotein N-acetylglucosaminyltransferase IVa; GlcNAc-T IVa; GnT-IVa; N-acetylglucosaminyltransferase IVa; UDP-N-acetylglucosamine: alpha-1,3-D-mannoside beta-1,4-N-acetylglucosaminyltransferase IVa.", 
            "tax_name": "Homo sapiens", 
            "gene": "MGAT4A; GnT-4a; GNT-IV; GNT-IVA.", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9UQ53-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9UQ53", 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase B; 2.4.1.145; N-glycosyl-oligosaccharide-glycoprotein N-acetylglucosaminyltransferase IVb; GlcNAc-T IVb; GnT-IVb; N-acetylglucosaminyltransferase IVb; UDP-N-acetylglucosamine: alpha-1,3-D-mannoside beta-1,4-N-acetylglucosaminyltransferase IVb.", 
            "tax_name": "Homo sapiens", 
            "gene": "MGAT4B; PRO1927; GNT-IV; GNT-IVB.", 
            "tax_id": 9606
        }, 
        {
            "uniprot_canonical_ac": "Q9UBM8-1", 
            "gene_link": "https://www.uniprot.org/uniprot/Q9UBM8", 
            "protein_name": "Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase C; 2.4.1.145; N-glycosyl-oligosaccharide-glycoprotein N-acetylglucosaminyltransferase IVc; GlcNAc-T IVc; GnT-IVc; N-acetylglucosaminyltransferase IVc; N-acetylglucosaminyltransferase IV homolog; hGnT-IV-H; UDP-N-acetylglucosamine: alpha-1,3-D-mannoside beta-1,4-N-acetylglucosaminyltransferase IVc.", 
            "tax_name": "Homo sapiens", 
            "gene": "MGAT4C; GNTIVH; HGNT-IV-H.", 
            "tax_id": 9606
        }
    ], 
    "crossref": [
        {
            "url": "https://glyconnect.expasy.org/browser/structures/2563", 
            "id": "2563", 
            "database": "GlyConnect"
        }, 
        {
            "url": "http://www.glycosciences.de/database/start.php?action=explore_linucsid&linucsid=24752", 
            "id": "24752", 
            "database": "Glycosciences.de"
        }, 
        {
            "url": "http://www.unicarbkb.org/structure/4211", 
            "id": "4211", 
            "database": "UniCarbKB"
        }
    ], 
    "glycoprotein": [
        {
            "residue": "Asn", 
            "uniprot_canonical_ac": "P07602-1", 
            "evidence": [
                {
                    "url": "http://unicarbkb.org/proteinsummary/P07602/annotated", 
                    "id": "P07602", 
                    "database": "UniCarbKB"
                }, 
                {
                    "url": "https://www.ncbi.nlm.nih.gov/pubmed/8344278", 
                    "id": "8344278", 
                    "database": "PubMed"
                }, 
                {
                    "url": "https://www.ncbi.nlm.nih.gov/pubmed/2110822", 
                    "id": "2110822", 
                    "database": "PubMed"
                }, 
                {
                    "url": "https://www.ncbi.nlm.nih.gov/pubmed/11180632", 
                    "id": "11180632", 
                    "database": "PubMed"
                }
            ], 
            "protein_name": "Prosaposin; Sphingolipid activator protein 2; SAP-2; Sphingolipid activator protein 1; SAP-1; Cerebroside sulfate activator; CSAct; Dispersin; Proactivator polypeptide; GM1 activator; A1 activator; Protein A; Component C; Glucosylceramidase activator; Co-beta-glucosidase; Protein C.", 
            "position": 215, 
            "tax_name": "Homo sapiens", 
            "gene_name": "PSAP; SAP1; GLBA; SAP2.", 
            "tax_id": 9606
        }
    ], 
    "mass_pme": 1567.81, 
    "inchi_key": {}, 
    "publication": [
        {
            "title": "Structure of the asparagine-linked sugar chains of porcine kidney and human urine cerebroside sulfate activator protein.", 
            "url": "https://www.ncbi.nlm.nih.gov/pubmed/11180632", 
            "journal": "Journal of mass spectrometry : JMS", 
            "evidence": [
                {
                    "url": "http://www.unicarbkb.org/structure/4211", 
                    "id": "4211", 
                    "database": "UniCarbKB"
                }
            ], 
            "authors": "Faull KF, Johnson J, Kim MJ, To T, Whitelegge JP, Stevens RL, Fluharty CB, Fluharty AL", 
            "date": "2000", 
            "pmid": "11180632"
        }, 
        {
            "title": "Structural study of the oligosaccharide moieties of sphingolipid activator proteins, saposins A, C and D obtained from the spleen of a Gaucher patient.", 
            "url": "https://www.ncbi.nlm.nih.gov/pubmed/8344278", 
            "journal": "European journal of biochemistry", 
            "evidence": [
                {
                    "url": "http://www.unicarbkb.org/structure/4211", 
                    "id": "4211", 
                    "database": "UniCarbKB"
                }
            ], 
            "authors": "Ito K, Takahashi N, Takahashi A, Shimada I, Arata Y, O'Brien JS, Kishimoto Y", 
            "date": "1993", 
            "pmid": "8344278"
        }, 
        {
            "title": "Characteristics of asparagine-linked sugar chains of sphingolipid activator protein 1 purified from normal human liver and GM1 gangliosidosis (type 1) liver.", 
            "url": "https://www.ncbi.nlm.nih.gov/pubmed/2110822", 
            "journal": "Biochemistry", 
            "evidence": [
                {
                    "url": "http://www.unicarbkb.org/structure/4211", 
                    "id": "4211", 
                    "database": "UniCarbKB"
                }
            ], 
            "authors": "Yamashita K, Inui K, Totani K, Kochibe N, Furukawa M, Okada S", 
            "date": "1990", 
            "pmid": "2110822"
        }
    ], 
    "smiles_isomeric": "", 
    "residues": [
        {
            "attachedby": "rxn.Q921Q3-1", 
            "detachedby": "", 
            "id": "NA", 
            "parentid": "no_id", 
            "name": "N-glycan_core_b-D-GlcpNAc_A"
        }, 
        {
            "attachedby": "rxn.Q921Q3-1", 
            "detachedby": "", 
            "id": "NB", 
            "parentid": "NA", 
            "name": "N-glycan_core_b-D-GlcpNAc_B"
        }, 
        {
            "attachedby": "rxn.Q921Q3-1", 
            "detachedby": "", 
            "id": "NC", 
            "parentid": "NB", 
            "name": "N-glycan_core_b-D-Manp"
        }, 
        {
            "attachedby": "rxn.Q921Q3-1", 
            "detachedby": "", 
            "id": "N1", 
            "parentid": "NC", 
            "name": "N-glycan_a-D-Manp_1"
        }, 
        {
            "attachedby": "rxn.Q921Q3-1", 
            "detachedby": "", 
            "id": "N7", 
            "parentid": "N1", 
            "name": "N-glycan_b-D-GlcpNAc_7"
        }, 
        {
            "attachedby": "rxn.Q921Q3-1", 
            "detachedby": "", 
            "id": "N4", 
            "parentid": "NC", 
            "name": "N-glycan_a-D-Manp_4"
        }, 
        {
            "attachedby": "rxn.Q921Q3-1", 
            "detachedby": "", 
            "id": "N15", 
            "parentid": "NA", 
            "name": "N-glycan_a-L-Fucp_15"
        }
    ], 
    "composition": [
        {
            "residue": "other", 
            "count": 0, 
            "name": "Other"
        }, 
        {
            "residue": "dhex", 
            "count": 1, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/840", 
            "name": "6-Deoxy-Hexose", 
            "cid": "840"
        }, 
        {
            "residue": "hexa", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/610", 
            "name": "Hexuronic Acid", 
            "cid": "610"
        }, 
        {
            "residue": "neugc", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/440001", 
            "name": "N-Glycolyl-Neuraminic Acid", 
            "cid": "440001"
        }, 
        {
            "residue": "hex", 
            "count": 3, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/206", 
            "name": "Hexose", 
            "cid": "206"
        }, 
        {
            "residue": "hexnac", 
            "count": 3, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/899", 
            "name": "N-Acetylhexosamine", 
            "cid": "899"
        }, 
        {
            "residue": "hexn", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/739", 
            "name": "2-Amino-2-Deoxy-Hexose", 
            "cid": "739"
        }, 
        {
            "residue": "p", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1061", 
            "name": "Phosphate", 
            "cid": "1061"
        }, 
        {
            "residue": "s", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/1117", 
            "name": "Sulfate", 
            "cid": "1117"
        }, 
        {
            "residue": "neuac", 
            "count": 0, 
            "url": "https://pubchem.ncbi.nlm.nih.gov/compound/439197", 
            "name": "N-Acetyl-Neuraminic Acid", 
            "cid": "439197"
        }
    ], 
    "glycam": "DGlcpNAcb1-4DManpa1-3[DManpa1-6]DManpb1-4DGlcpNAcb1-4[LFucpa1-6]DGlcpNAcb1-", 
    "number_monosaccharides": 7, 
    "motifs": [
        {
            "id": "G00026MO", 
            "name": "N-Glycan core basic"
        }
    ], 
    "inchi": ""
}